D. Bates, M. Maechler, B. Bolker, and S. Walker, lme4: Linear mixed-effect models using 350 Eigen and S4. R package version 1.0-5, pp.4-351, 2013.

Y. Benjamini and Y. Hochberg, Controlling the false discovery rate: a practical and 352 powerful approach to multiple testing, J. R. Stat. Soc. Ser. B. Methodol, vol.57, pp.289-300, 1995.

A. Bockelmann and R. Neuhaus, from a high-stress habitat in a European salt marsh, Journal of Ecology, vol.113, issue.3, pp.503-513, 1999.
DOI : 10.1046/j.1365-2745.1999.00368.x

P. Cardoso, S. Pekár, R. Jocqué, and J. A. Coddington, Global patterns of guild 356 composition and functional diversity of spiders, PLoS One, vol.6, p.357, 2011.

S. Villéger and N. Mouquet, Defining and measuring ecological specialization, J, p.359, 2010.

V. Devictor, R. Julliard, J. Clavel, F. Jiguet, A. Lee et al., Functional biotic homogenization of bird communities in disturbed landscapes, Global Ecology and Biogeography, vol.70, issue.2, pp.252-261, 2008.
DOI : 10.1038/35102054

V. Devictor, D. Mouillot, C. Meynard, F. Jiguet, W. Thuiller et al., Spatial 364 mismatch and congruence between taxonomic, phylogenetic and functional diversity: 365 the need for integrative conservation strategies in a changing world, Ecol. Lett, vol.13, issue.366, pp.1030-1070, 2010.

V. Devictor and A. Robert, Measuring community responses to large-scale disturbance in conservation biogeography, Diversity and Distributions, vol.11, issue.1, pp.122-130, 2009.
DOI : 10.1111/j.1472-4642.2008.00510.x

A. Doxa, Y. Bas, M. L. Paracchini, P. Pointereau, J. Terres et al., Low-intensity agriculture increases farmland bird abundances in France, Journal of Applied Ecology, vol.5, issue.6, pp.1348-1356, 2010.
DOI : 10.1111/j.1365-2664.2010.01869.x

H. Ford, A. Garbutt, L. Jones, and D. L. Jones, Grazing management in saltmarsh ecosystems drives invertebrate diversity, abundance and functional group structure, Insect Conservation and Diversity, vol.6, issue.4, pp.189-200, 2012.
DOI : 10.1111/j.1752-4598.2012.00202.x

T. Hothorn, F. Bretz, and P. Westfall, Simultaneous Inference in General Parametric Models, Biometrical Journal, vol.16, issue.3, pp.346-363, 2008.
DOI : 10.1002/bimj.200810425

R. Julliard, J. Clavel, V. Devictor, F. Jiguet, and D. Couvet, Spatial segregation of specialists and generalists in bird communities, Ecology Letters, vol.135, issue.11, pp.1237-1281, 2006.
DOI : 10.1086/285702

URL : https://hal.archives-ouvertes.fr/hal-00113560

E. Laliberté and B. Shipley, FD: measuring functional diversity from multiple traits, and 381 other tools for functional ecology. R package version 1, pp.0-11, 2011.

B. Leroy, Rarity: Calculation of rarity indices for species and assemblages of species. R 383 package version 1.2-1, p.384, 2013.

B. Leroy, A. Canard, and F. Ysnel, Integrating multiple scales in rarity assessments of invertebrate taxa, Diversity and Distributions, vol.24, issue.7, pp.794-803, 2013.
DOI : 10.1111/ddi.12040

B. Leroy, J. Pétillon, R. Gallon, A. Canard, and F. Ysnel, Improving occurrence-based 387 rarity metrics in conservation studies by including multiple rarity cut-off points, pp.159-168, 2012.

R. Levins, Evolution in Changing Environments: Some Theoretical Explorations, 1968.

P. Marc, A. Canard, and F. Ysnel, Spiders (Araneae) useful for pest limitation and bioindication, Agriculture, Ecosystems & Environment, vol.74, issue.1-3, pp.229-273, 1999.
DOI : 10.1016/S0167-8809(99)00038-9

C. N. Meynard, V. Devictor, D. Mouillot, W. Thuiller, F. Jiguet et al., Beyond 394 taxonomic diversity patterns: how do ?, ? and ? components of bird functional and 395 phylogenetic diversity respond to environmental gradients across France? Glob. Ecol. 396 Biogeogr, pp.893-903, 2011.

M. Kulbicki, S. Lavergne, S. Lavorel, N. Mouquet, C. E. Paine et al., Rare species support vulnerable functions in high-diversity ecosystems
URL : https://hal.archives-ouvertes.fr/hal-00912633

C. Penone, C. Kerbiriou, J. Julien, R. Julliard, N. Machon et al., Urbanisation effect on Orthoptera: which scale matters?, Insect Conservation and Diversity, vol.101, issue.3, pp.319-403, 2013.
DOI : 10.1111/j.1752-4598.2012.00217.x

J. Pétillon, A. Georges, A. Canard, J. Lefeuvre, J. P. Bakker et al., Influence of abiotic factors on spider and ground beetle communities in different salt-marsh systems, Basic and Applied Ecology, vol.9, issue.6, pp.743-751, 2008.
DOI : 10.1016/j.baae.2007.08.007

J. Pétillon, A. Georges, A. Canard, and F. Ysnel, Impact of cutting and sheep grazing on 408 ground-active spiders and carabids in intertidal salt marshes (Western France). Anim. 409, Biodivers. Conserv, vol.30, pp.201-209, 2007.

J. Pétillon, K. Lambeets, B. Ract-madoux, P. Vernon, and D. Renault, Saline stress 411 tolerance partly matches with habitat preference in ground-living wolf spiders, pp.165-172, 2011.

J. Pétillon, F. Ysnel, A. Canard, and J. Lefeuvre, Impact of an invasive plant (Elymus athericus) on the conservation value of tidal salt marshes in western France and implications for management: Responses of spider populations, Biological Conservation, vol.126, issue.1, pp.103-416, 2005.
DOI : 10.1016/j.biocon.2005.05.003

S. Prieto-benítez and M. Méndez, Effects of land management on the abundance and richness of spiders (Araneae): A meta-analysis, Biological Conservation, vol.144, issue.2, pp.683-691, 2011.
DOI : 10.1016/j.biocon.2010.11.024

C. Rickert, A. Fichtner, R. Van-klink, and J. P. Bakker, ??- and ??-diversity in moth communities in salt marshes is driven by grazing management, Biological Conservation, vol.146, issue.1, pp.24-423
DOI : 10.1016/j.biocon.2011.11.024

O. J. Schmitz, Effects of predator functional diversity on grassland ecosystem function, Ecology, vol.90, issue.9, pp.2339-2345, 2009.
DOI : 10.1111/j.1461-0248.2005.00850.x

A. Strecker, J. Olden, J. B. Whittier, and C. P. Paukert, Defining conservation priorities for 427 freshwater fishes according to taxonomic, functional, and phylogenetic diversity, Ecol, p.428, 2011.

C. J. Topping and K. D. Sunderland, Limitations to the use of pitfall traps in ecological 430 studies exemplified by a study of spiders in a field of winter wheat, J. Appl. Ecol, vol.29, pp.431-485, 1992.

G. W. Uetz, J. Halaj, and A. B. Cady, Guild structure of spiders in major crops, J. Arachnol, vol.433, pp.27-270, 1999.

R. Van-klink, C. Rickert, R. Vermeulen, O. Vorst, M. F. Wallisdevries et al., Grazed vegetation mosaics do not maximize arthropod diversity: Evidence from salt marshes, Biological Conservation, vol.164, pp.150-157, 2013.
DOI : 10.1016/j.biocon.2013.04.023

R. M. Veeneklaas, K. S. Dijkema, N. Hecker, and J. P. Bakker, on natural salt marshes, Applied Vegetation Science, vol.23, issue.2, pp.205-216, 2012.
DOI : 10.1111/j.1654-109X.2012.01228.x

S. Villéger, N. W. Mason, and D. Mouillot, NEW MULTIDIMENSIONAL FUNCTIONAL DIVERSITY INDICES FOR A MULTIFACETED FRAMEWORK IN FUNCTIONAL ECOLOGY, Ecology, vol.89, issue.8, pp.2290-2301, 2008.
DOI : 10.1111/j.1461-0248.2005.00850.x