J. Altmann, Observational Study of Behavior: Sampling Methods, Behaviour, vol.49, issue.3, pp.227-267, 1974.
DOI : 10.1163/156853974X00534

C. A. Anderson and B. J. Bushman, Human Aggression, Annual Review of Psychology, vol.53, issue.1, pp.27-51, 2002.
DOI : 10.1146/annurev.psych.53.100901.135231

D. Antonacci, I. Norscia, and E. Palagi, Stranger to Familiar: Wild Strepsirhines Manage Xenophobia by Playing, PLoS ONE, vol.56, issue.3, 2010.
DOI : 10.1371/journal.pone.0013218.s003

J. D. Baldwin, Behavior in infancy: Exploration and play, Comparative Primate Biology, vol.2, pp.295-326, 1986.

P. Bateson and M. Young, Separation from the mother and the development of play in cats, Animal Behaviour, vol.29, issue.1, pp.173-180, 1981.
DOI : 10.1016/S0003-3472(81)80163-7

P. Bateson, M. Mendl, and J. Feaver, Play in the domestic cat is enhanced by rationing of the mother during lactation, Animal Behaviour, vol.40, issue.3, pp.514-525, 1990.
DOI : 10.1016/S0003-3472(05)80532-9

M. Bekoff, Play Signals as Punctuation: the Structure of Social Play in Canids, Behaviour, vol.132, issue.5, pp.419-429, 1995.
DOI : 10.1163/156853995X00649

M. Bekoff and J. A. Byers, A critical reanalysis of the ontogeny of mammalian social and locomotor play: An ethological hornet's nest Behavioral development: The Bielefeld interdisciplinary project, pp.296-337, 1981.

M. Bekoff and J. A. Byers, Animal play ? Evolutionary, comparative and ecological perspectives, 1998.

H. Benhajali, M. A. Richard-yris, M. Leroux, M. Ezzaouia, F. Charfi et al., A note on the time budget and social behaviour of densely housed horses, Applied Animal Behaviour Science, vol.112, issue.1-2, pp.196-200, 2008.
DOI : 10.1016/j.applanim.2007.08.007
URL : https://hal.archives-ouvertes.fr/hal-01319146

I. S. Bernstein and W. A. Mason, Group formation by rhesus monkeys, Animal Behaviour, vol.11, issue.1, pp.28-31, 1963.
DOI : 10.1016/0003-3472(63)90004-6

M. Biben and M. Champoux, Play and stress: Cortisol as a negative correlate of play in Saimiri, Play and culture studies, pp.191-208, 1999.

D. T. Blumstein, L. K. Chung, and J. E. Smith, Early play may predict later dominance relationships in yellow-bellied marmots (Marmota flaviventris), Proceedings of the Royal Society B: Biological Sciences, vol.440, issue.7088, 2013.
DOI : 10.1038/nature04540

A. Boissy, G. Manteuffel, M. B. Jensen, R. O. Moe, B. Spruijt et al., Assessment of positive emotions in animals to improve their welfare, Physiology & Behavior, vol.92, issue.3, pp.375-397, 2007.
DOI : 10.1016/j.physbeh.2007.02.003

M. Bourjade, A. De-boyer-des-roches, and M. Hausberger, Adult-Young Ratio, a Major Factor Regulating Social Behaviour of Young: A Horse Study, PLoS ONE, vol.20, issue.5, p.4888, 2009.
DOI : 10.1371/journal.pone.0004888.t001
URL : https://hal.archives-ouvertes.fr/hal-01237436

M. Bourjade, M. Moulinot, M. A. Richard-yris, and M. Hausberger, Could adults be used to improve social skills of young horses, Equus caballus? Developmental Psychobiology, pp.408-417, 2008.

L. Brent and Y. Viera, Social behavior of captive indochinese and insular long-tailed macaques following transfer to new facility, International Journal of Primatology, vol.23, issue.1, pp.147-159, 2002.
DOI : 10.1023/A:1013206125884

S. Budiansky, The nature of horses, 1997.

G. M. Burghardt, The genesis of animal play: Testing the limits, 2005.

C. C. Burn, T. L. Dennison, and H. R. Whay, Relationships between behaviour and health in working horses, donkeys, and mules in developing countries, Applied Animal Behaviour Science, vol.126, issue.3-4, pp.109-118, 2010.
DOI : 10.1016/j.applanim.2010.06.007

J. A. Byers, American pronghorn. Social adaptation and the ghosts of predators past, 1997.

J. A. Byers, Biological effects of locomotor play: Getting into shape or something more specific Animal play ? Evolutionary, comparative and ecological perspectives, pp.205-220, 1998.

J. A. Byers and C. Walker, Refining the Motor Training Hypothesis for the Evolution of Play, The American Naturalist, vol.146, issue.1, pp.25-40, 1995.
DOI : 10.1086/285785

E. Z. Cameron, W. L. Linklater, K. J. Stafford, and E. O. Minot, Maternal investment results in better foal condition through increased play behaviour in horses, Animal Behaviour, vol.76, issue.5, pp.1511-1518, 2008.
DOI : 10.1016/j.anbehav.2008.07.009

S. Camus, C. Blois-heulin, Q. Li, M. Hausberger, and E. Bezard, Behavioural Profiles in Captive-Bred Cynomolgus Macaques: Towards Monkey Models of Mental Disorders?, PLoS ONE, vol.53, issue.4, 2013.
DOI : 10.1371/journal.pone.0062141.s002
URL : https://hal.archives-ouvertes.fr/hal-01021752

S. Camus, C. Rochais, C. Blois-heulin, Q. Li, M. Hausberger et al., Birth Origin Differentially Affects Depressive-Like Behaviours: Are Captive-Born Cynomolgus Monkeys More Vulnerable to Depression than Their Wild-Born Counterparts?, PLoS ONE, vol.282, issue.7, 2013.
DOI : 10.1371/journal.pone.0067711.s004
URL : https://hal.archives-ouvertes.fr/hal-01052723

S. Camus, C. Rochais, C. Blois-heulin, Q. Li, M. Hausberger et al., Depressive-like behavioural profiles in captive-bred single-and socially-housed rhesus and cynomolgus macaques: A species comparison, Frontiers in Behavioral Neuroscience, vol.8, pp.1-15, 2014.

H. A. Carr, A Statistical Study of Education in the West, The American Journal of Psychology, vol.13, issue.4, pp.1-47, 1902.
DOI : 10.2307/1412451

J. W. Christensen, J. Ladewig, E. Søndergaard, and J. Malmkvist, Effects of individual versus group stabling on social behaviour in domestic stallions, Applied Animal Behaviour Science, vol.75, issue.3, pp.233-248, 2002.
DOI : 10.1016/S0168-1591(01)00196-4

F. Ciani, S. Dall-'olio, R. Tanyon, and E. Palagi, Social tolerance and adult play in macaque societies: a comparison with different human cultures, Animal Behaviour, vol.84, issue.6, pp.1313-1322, 2012.
DOI : 10.1016/j.anbehav.2012.09.002

R. Clubb and G. Mason, A review of the welfare of zoo elephants in Europe, 2002.

C. L. Coe, G. R. Lubach, W. B. Ershler, and R. G. Klopp, Influence of early rearing on lymphocyte proliferation responses in juvenile rhesus monkeys, Brain, Behavior, and Immunity, vol.3, issue.1, pp.47-60, 1989.
DOI : 10.1016/0889-1591(89)90005-6

R. P. Coppinger, J. I. Glendinning, E. Torop, C. Matthay, M. Sutherland et al., Degree of Behavioral Neoteny Differentiates Canid Polymorphs, Ethology, vol.10, issue.2, 1987.
DOI : 10.1111/j.1439-0310.1987.tb00645.x

R. Coppinger and C. K. Smith, A model for understanding the evolution of mammalian behavior Current mammalogy, pp.335-374, 1989.

C. Corino, R. Rossi, M. Musella, S. Cannata, and G. Pastorelli, Growth performance and oxidative status in piglets supplemented with verbascoside and teupolioside, Italian Journal of Animal Science, vol.6, issue.1s, pp.292-294, 2007.
DOI : 10.4081/ijas.2007.1s.292

C. M. Crockett, C. L. Bowers, M. Shimoji, M. Leu, D. M. Bowden et al., Behavioral responses of longtailed macaques to different cage sizes and common laboratory experiences., Journal of Comparative Psychology, vol.109, issue.4, pp.368-383, 1995.
DOI : 10.1037/0735-7036.109.4.368

S. L. Crowell-davis, K. A. Houpt, and L. Kane, Play development in Welsh pony (Equus caballus) foals, Applied Animal Behaviour Science, vol.18, issue.2, pp.119-131, 1987.
DOI : 10.1016/0168-1591(87)90186-9

R. K. Davenport and E. W. Menzel, Stereotyped Behavior of the Infant Chimpanzee, Archives of General Psychiatry, vol.8, issue.1, pp.99-104, 1963.
DOI : 10.1001/archpsyc.1963.01720070101013

J. E. Day, H. A. De-weerd, and S. A. Edwards, The effect of varying lengths of straw bedding on the behaviour of growing pigs, Applied Animal Behaviour Science, vol.109, issue.2-4, pp.249-260, 2008.
DOI : 10.1016/j.applanim.2007.02.006

D. Moffarts, B. Kirschvink, N. Pincemail, J. Lekeux, and P. , Physiological and pathological impact of oxidative stress in the equine species, pp.1-9, 2005.

B. J. Devinney, C. M. Berman, and K. L. Rasmussen, Individual differences in response to sibling birth among free-ranging yearling rhesus monkeys (Macaca mulatta) on Cayo Santiago, Behaviour, vol.140, issue.7, pp.899-924, 2003.
DOI : 10.1163/156853903770238373

A. Diener, Behaviour analysis of polecats ferrets during social play, pp.179-197, 1985.

T. M. Donaldson, R. C. Newberry, M. Spinka, and S. Cloutier, Effects of early play experience on play behaviour of piglets after weaning, Applied Animal Behaviour Science, vol.79, issue.3, pp.221-231, 2002.
DOI : 10.1016/S0168-1591(02)00138-7

S. Dudink, H. Simonse, I. Marks, F. H. De-jonge, and B. M. Spruijt, Announcing the arrival of enrichment increases play behaviour and reduces weaning-stress-induced behaviours of piglets directly after weaning, Applied Animal Behaviour Science, vol.101, issue.1-2, pp.86-101, 2006.
DOI : 10.1016/j.applanim.2005.12.008

V. Durier, S. Henry, C. Sankey, J. Sizun, and M. Hausberber, Locomotor Inhibition in Adult Horses Faced to Stressors: A Single Postpartum Experience May be Enough!, Frontiers in Psychology, vol.3, 2012.
DOI : 10.3389/fpsyg.2012.00442
URL : https://hal.archives-ouvertes.fr/hal-01020434

R. Fagen and F. Fagen, Juvenile survival and benefits of play behavior in brown bears, Ursus arctos, Evolutionary Ecology Research, vol.6, pp.89-102, 2004.

C. Feh, Relationships and communication in socially natural horse herd The domestic horse, the origins, development and management of its behaviour, pp.83-93, 2005.

J. D. Feist and D. R. Mccullough, Behaviour patterns and communication in feral horses, Zeitschrift für Tierzuchtung und Zuchtungsbiologie, pp.337-373, 1976.

G. F. Fisler, Nonbreeding Activities of Three Adult Males in a Band of Free-Ranging Rhesus Monkeys, Journal of Mammalogy, vol.48, issue.1, pp.70-78
DOI : 10.2307/1378171

B. P. Fonseca, A. L. Alves, J. L. Nicoletti, A. Thomassian, C. A. Hussni et al., Thermography and ultrasonography in back pain diagnosis of equine athletes, Journal of Equine Veterinary Science, vol.26, issue.11, pp.507-516, 2006.
DOI : 10.1016/j.jevs.2006.09.007

A. F. Fraser and D. M. Broom, Farm animals and welfare, 1990.

R. O. Frost, G. Steketee, and K. A. Green, Cognitive and Behavioral Treatment of Compulsive Hoarding, Brief Treatment and Crisis Intervention, vol.3, issue.3, pp.323-338, 2003.
DOI : 10.1093/brief-treatment/mhg024

C. Fureix, H. Benhajali, S. Henry, A. Bruchet, A. Prunier et al., Plasma cortisol and faecal cortisol metabolites concentrations in stereotypic and non-stereotypic horses: do stereotypic horses cope better with poor environmental conditions?, BMC Veterinary Research, vol.9, issue.1, pp.10-1186, 2013.
DOI : 10.1046/j.1439-0442.1999.00256.x
URL : https://hal.archives-ouvertes.fr/hal-01019939

C. Fureix, A. Gorecka-bruzda, E. Gautier, and M. Hausberger, Co-occurrence of yawning and stereotypic behaviour in horses Equus caballus, 2011.

C. Fureix, P. Jégo, S. Henry, L. Lansade, and M. Hausberger, Towards an Ethological Animal Model of Depression? A Study on Horses, PLoS ONE, vol.427, issue.6, 2012.
DOI : 10.1371/journal.pone.0039280.t001
URL : https://hal.archives-ouvertes.fr/hal-01020448

C. Fureix, P. Jégo, C. Sankey, and M. Hausberger, How horses (Equus caballus) see the world: humans as significant ???objects???, Animal Cognition, vol.40, issue.1, pp.643-654, 2009.
DOI : 10.1007/s10071-009-0223-2
URL : https://hal.archives-ouvertes.fr/hal-01319138

C. Fureix, H. Menguy, and M. Hausberger, Partners with Bad Temper: Reject or Cure? A Study of Chronic Pain and Aggression in Horses, PLoS ONE, vol.112, issue.4, p.12434, 2010.
DOI : 10.1371/journal.pone.0012434.g005
URL : https://hal.archives-ouvertes.fr/hal-01019967

K. L. Graham, Virtually playful: Do massively multiplayer online role-playing games meet behavioral criteria of play?, American Journal of Play, vol.3, pp.106-125, 2010.

K. L. Graham, Coevolutionary relationship between striatum size and social play in nonhuman primates, American Journal of Primatology, vol.23, issue.4, pp.314-322, 2011.
DOI : 10.1002/ajp.20898

K. L. Gross and . Baldwin, The play of animals, 1898.

R. F. Guerra, E. Takase, and C. R. Nunes, Play fighting of juvenile golden hamsters (Mesocricetus auratus): effects of two types of social deprivation and days of testing, Behavioural Processes, vol.47, issue.3, pp.139-151, 1999.
DOI : 10.1016/S0376-6357(99)00058-3

S. L. Hall and J. W. Bradshaw, The influence of hunger on object play by adult domestic cats, Applied Animal Behaviour Science, vol.58, issue.1-2, pp.143-150, 1998.
DOI : 10.1016/S0168-1591(97)00136-6

C. Hall, D. Goodwin, C. Heleski, H. Randle, and N. Waran, Is There Evidence of Learned Helplessness in Horses?, Journal of Applied Animal Welfare Science, vol.40, issue.3, pp.249-266, 2008.
DOI : 10.1080/10888700802101130

M. Hausberger, C. Fureix, M. Bourjade, S. Wessel-robert, and M. A. Richard-yris, On the significance of adult play: what does social play tell us about adult horse welfare?, Naturwissenschaften, vol.43, issue.5, pp.291-302, 2012.
DOI : 10.1007/s00114-012-0902-8
URL : https://hal.archives-ouvertes.fr/hal-01021858

M. Hausberger, E. Gautier, V. Biquand, C. Lunel, and P. Jégo, Could Work Be a Source of Behavioural Disorders? A Study in Horses, PLoS ONE, vol.13, issue.2, 2009.
DOI : 10.1371/journal.pone.0007625.s001
URL : https://hal.archives-ouvertes.fr/hal-01319141

M. Hausberger, S. Henry, C. Larose, and M. A. Richard-yris, First suckling: A crucial event for mother-young attachment? An experimental study in horses (Equus caballus)., Journal of Comparative Psychology, vol.121, issue.1, pp.109-112, 2007.
DOI : 10.1037/0735-7036.121.1.109
URL : https://hal.archives-ouvertes.fr/hal-01319147

M. Hausberger, C. Muller, and C. Lunel, Does Work Affect Personality? A Study in Horses, PLoS ONE, vol.22, issue.10, 2011.
DOI : 10.1371/journal.pone.0014659.s004
URL : https://hal.archives-ouvertes.fr/hal-01022090

M. Hausberger, H. Roche, S. Henry, and E. K. Visser, A review of the human???horse relationship, Applied Animal Behaviour Science, vol.109, issue.1, pp.1-24, 2008.
DOI : 10.1016/j.applanim.2007.04.015
URL : https://hal.archives-ouvertes.fr/hal-01319144

S. D. Held and M. Spinka, Animal play and animal welfare, Animal Behaviour, vol.81, issue.5, pp.891-899, 2011.
DOI : 10.1016/j.anbehav.2011.01.007

S. Henry, M. A. Richard-yris, S. Tordjman, and M. Hausberger, Neonatal Handling Affects Durably Bonding and Social Development, PLoS ONE, vol.49, issue.4, p.5216, 2009.
DOI : 10.1371/journal.pone.0005216.t001
URL : https://hal.archives-ouvertes.fr/hal-01319140

S. Henry, A. J. Zanella, C. Sankey, M. A. Richard-yris, A. Marko et al., Adults may be used to alleviate weaning stress in domestic foals (Equus caballus), Physiology & Behavior, vol.106, issue.4, pp.428-438, 2012.
DOI : 10.1016/j.physbeh.2012.02.025
URL : https://hal.archives-ouvertes.fr/hal-01020422

B. T. Himmler, R. Stryjek, K. Modlinska, S. M. Derksen, W. Pisula et al., How domestication modulates play behavior: A comparative analysis between wild rats and a laboratory strain of Rattus norvegicus., Journal of Comparative Psychology, vol.127, issue.4, 2013.
DOI : 10.1037/a0032187

R. Hoffmann, On the development of social behavior in immature males of a feral horse population (Equusprzewalskii caballus), Zeitschrift für Saugetierkunde International Journal of Mammalian Biology, vol.50, pp.302-314, 1985.

L. B. Jeffcott, Disorders of the thoracolumbar spine of the horse - a survey of 443 cases, Equine Veterinary Journal, vol.59, issue.4, pp.197-210, 1980.
DOI : 10.1111/j.2042-3306.1980.tb03427.x

M. B. Jensen and R. Kyhn, Play behaviour in group-housed dairy calves, the effect of space allowance, Applied Animal Behaviour Science, vol.67, issue.1-2, pp.35-46, 2000.
DOI : 10.1016/S0168-1591(99)00113-6

J. A. Joireman, C. S. Fick, and J. W. Anderson, Sensation seeking and involvement in chess, Personality and Individual Differences, vol.32, issue.3, pp.509-515, 2002.
DOI : 10.1016/S0191-8869(01)00052-6

B. N. Khan, Impact of captivity on social behaviour of chimpanzee (Pan troglodytes), The Journal of Animal & Plant Sciences, vol.23, pp.779-785, 2013.

Z. A. Klein, W. A. Padow, and R. D. Romeo, The effects of stress on play and home cage behaviors in adolescent male rats, Developmental Psychobiology, vol.33, pp.62-70, 2010.
DOI : 10.1002/dev.20413

L. Lansade, G. Pichard, and M. Leconte, Sensory sensitivities: Components of a horse's temperament dimension, Applied Animal Behaviour Science, vol.114, issue.3-4, pp.534-553, 2008.
DOI : 10.1016/j.applanim.2008.02.012

A. Lemasson, C. Blois-heulin, R. Jubin, and M. Hausberger, Female social relationships in a captive group of Campbell's monkeys (Cercopithecus campbelli campbelli), American Journal of Primatology, vol.140, issue.12, pp.1161-1170, 2006.
DOI : 10.1002/ajp.20315

A. Lemasson, J. P. Gautier, and M. Hausberger, A brief note on the effects of the removal of individuals on social behaviour in a captive group of campbell's monkeys (Cercopithecus campbelli campbelli): a case study, Applied Animal Behaviour Science, vol.91, issue.3-4, pp.289-296, 2005.
DOI : 10.1016/j.applanim.2004.10.011

C. Lesimple, C. Fureix, V. Biquand, and M. Hausberger, Comparison of clinical examinations of back disorders and humans??? evaluation of back pain in riding school horses, BMC Veterinary Research, vol.9, issue.1, p.209, 2013.
DOI : 10.1016/j.physbeh.2007.02.003
URL : https://hal.archives-ouvertes.fr/hal-01019934

C. Lesimple, C. Fureix, L. Scolan, N. Richard-yris, M. A. Hausberger et al., Housing conditions and breed are associated with emotionality and cognitive abilities in riding school horses, Applied Animal Behaviour Science, vol.129, issue.2-4, pp.92-99, 2011.
DOI : 10.1016/j.applanim.2010.11.005
URL : https://hal.archives-ouvertes.fr/hal-01022148

A. Loranca, C. Terrero, and M. L. Salas, Development of Play Behavior in Neonatally Undernourished Rats, Physiology & Behavior, vol.66, issue.1, pp.3-10, 1999.
DOI : 10.1016/S0031-9384(98)00235-2

G. Mancini and E. Palagi, Play and social dynamics in a captive herd of gelada baboons (Theropithecus gelada), Behavioural Processes, vol.82, issue.3, pp.286-292, 2009.
DOI : 10.1016/j.beproc.2009.07.007

R. A. Martin, Is Laughter the Best Medicine? Humor, Laughter, and Physical Health, Current Directions in Psychological Science, vol.11, issue.6, pp.216-220, 2002.
DOI : 10.1037/0033-295X.96.2.234

G. J. Mason, Stereotypies and suffering, Behavioural Processes, vol.25, issue.2-3, pp.103-115, 1991.
DOI : 10.1016/0376-6357(91)90013-P

G. J. Mason, Early weaning enhances the later development of stereotypy in minks, Proceedings of the 30th International Congress of the ISAE, p.16, 1996.

G. J. Mason and P. C. Green, The effects of social restriction on the behavior of rhesus monkeys: IV. Responses to a novel environment and to an alien species., Journal of Comparative and Physiological Psychology, vol.55, issue.3, pp.363-368, 1962.
DOI : 10.1037/h0044807

G. J. Mason and N. R. Latham, Can't stop, won't stop: Is stereotypy a reliable animal welfare indicator? Animal Welfare, pp.57-69, 2004.

G. J. Mason and J. Rushen, Stereotypic behavior in captive animals: Fundamentals and applications for welfare, 2006.

M. Mcdonnell and A. Poulin, Equid play ethogram, Applied Animal Behaviour Science, vol.78, issue.2-4, pp.263-290, 2002.
DOI : 10.1016/S0168-1591(02)00112-0

P. D. Mcgreevy, P. J. Cripps, N. P. French, L. E. Green, and C. J. Nicol, Management factors associated with stereotypic and redirected behaviour in the Thoroughbred horse, Equine Veterinary Journal, vol.14, issue.2, pp.86-91, 1995.
DOI : 10.1111/j.2042-3306.1995.tb03041.x

D. Mendoza-granados and V. Sommer, Play in chimpanzees of the Arnhem Zoo: Self-serving compromises, Primates, vol.14, issue.1, pp.36-57, 1995.
DOI : 10.1007/BF02381915

D. S. Mills, Repetitive movement problems in the horse The domestic horse: The origins, development and management of its behaviour, pp.212-227, 2005.

E. M. Mintline, S. L. Wood, A. M. De-passillé, J. Rushen, and C. B. Tucker, Assessing calf play behavior in an arena test, Applied Animal Behaviour Science, vol.141, issue.3-4, pp.101-107, 2012.
DOI : 10.1016/j.applanim.2012.08.006

P. Mormède, S. Adanson, B. Aupérin, B. Beerda, D. Guémené et al., Exploration of the hypothalamic???pituitary???adrenal function as a tool to evaluate animal welfare, Physiology & Behavior, vol.92, issue.3, pp.317-339, 2007.
DOI : 10.1016/j.physbeh.2006.12.003

R. C. Newberry and D. G. Wood-gush, ABSTRACT, Animal Production, vol.13, issue.01, pp.103-109, 1988.
DOI : 10.1016/0304-3762(76)90067-5

L. Normansell and J. Panksepp, Effects of morphine and naloxone on play-rewarded spatial discrimination in juvenile rats, Developmental Psychobiology, vol.95, issue.1, pp.75-83, 1990.
DOI : 10.1002/dev.420230108

I. Norscia and E. Palagi, When play is a family business: adult play, hierarchy, and possible stress reduction in common marmosets, Primates, vol.1, issue.2, pp.101-104, 2011.
DOI : 10.1007/s10329-010-0228-0

M. A. Novak, J. S. Meyer, C. Lutz, and S. Tiefenbacher, Deprived environments: developmental insights from primatology., Stereotypic animal behaviour: Fundamentals and applications to welfare, pp.153-189, 2006.
DOI : 10.1079/9780851990040.0153

S. Nunes, E. M. Muecke, J. A. Anthony, and A. S. Batterbee, Endocrine and Energetic Mediation of Play Behavior in Free-Living Belding's Ground Squirrels, Hormones and Behavior, vol.36, issue.2, pp.153-165, 1999.
DOI : 10.1006/hbeh.1999.1538

A. F. Oliveira, A. O. Rossi, L. F. Silva, M. C. Lau, and R. E. Barreto, Play behaviour in nonhuman animals and the animal welfare issue, Journal of Ethology, vol.75, issue.1, pp.1-5, 2010.
DOI : 10.1007/s10164-009-0167-7

O. Neill-wagner and P. L. , When trying to get your monkeys to behave, try perches, Touch, vol.1, pp.6-8, 1994.

N. W. Owens, A comparison of aggressive play and aggression in free-living baboons, Papio anubis, Animal Behaviour, vol.23, pp.755-465, 1975.
DOI : 10.1016/0003-3472(75)90103-7

E. Palagi, Play at work: Revisiting data focusing on chimpanzees (Pan troglodytes), Journal of Anthropological Sciences, vol.85, pp.153-164, 2007.

E. Palagi, G. Cordoni, and S. M. Borgognini, ), Ethology, vol.109, issue.12, pp.949-962, 2004.
DOI : 10.1111/j.1439-0310.2004.01035.x

E. Palagi, T. Paoli, and S. Borgognini-tarli, Short-Term Benefits of Play Behavior and Conflict Prevention in Pan paniscus, International Journal of Primatology, vol.109, issue.5, pp.1257-1270, 2006.
DOI : 10.1007/s10764-006-9071-y

J. Panksepp and W. W. Beatty, Social deprivation and play in rats, Behavioral and Neural Biology, vol.30, issue.2, pp.197-206, 1980.
DOI : 10.1016/S0163-1047(80)91077-8

S. M. Pellis and A. N. Iwaniuk, The Problem of Adult Play Fighting: A Comparative Analysis of Play and Courtship in Primates, Ethology, vol.123, issue.9, pp.783-806, 1999.
DOI : 10.1002/(SICI)1098-2337(1999)25:2<141::AID-AB6>3.3.CO;2-J

S. M. Pellis and M. M. Mckenna, Intrinsic and extrinsic influences on play fighting in rats: effects of dominance, partner's playfulness, temperament and neonatal exposure to testosterone propionate, Behavioural Brain Research, vol.50, issue.1-2, pp.135-145, 1992.
DOI : 10.1016/S0166-4328(05)80295-5

S. M. Pellis and V. C. Pellis, The playful brain: Venturing to the limits of neuroscience, 2009.

S. M. Pellis, V. C. Pellis, and H. C. Bell, The function of play in the development of the social brain, American Journal of Play, vol.2, pp.278-296, 2010.

S. M. Pellis, V. C. Pellis, and C. J. Reinhart, The evolution of social play Formative experiences: The interaction of caregiving, culture, and developmental psychobiology, 2010.

O. Petit and B. Thierry, Reconciliation in a group of Guinea baboons (Papio papio), Current primatology, pp.137-145, 1994.

W. Post and J. Baulu, Time budgets ofMacaca mulatta, Primates, vol.52, issue.1, pp.125-140, 1978.
DOI : 10.1007/BF02373230

L. Reamer, Z. Tooze, C. Coulson, and S. Semple, Correlates of self-directed and stereotypic behaviours in captive red-capped mangabey (Cercocebus torquatus torquatus) Applied Animal Behaviour Science, pp.68-74, 2010.

C. J. Reinhart, D. C. Mcintyre, G. Metz, and S. M. Pellis, Play fighting between kindling-prone (fast) and kindling-resistant (slow) rats., Journal of Comparative Psychology, vol.120, issue.1, pp.19-30, 2006.
DOI : 10.1037/0735-7036.120.1.19

C. Sherwin, Voluntary wheel running: a review and novel interpretation, Animal Behaviour, vol.56, issue.1, pp.11-27, 1998.
DOI : 10.1006/anbe.1998.0836

H. Sigurjonsdottir, M. C. Van-dierendonck, S. Snorrason, and A. G. Thorhallsdottir, Social relationships in a group of horses without a mature stallion, Behaviour, vol.140, issue.6, pp.783-804, 2003.
DOI : 10.1163/156853903322370670

S. M. Sivyi, C. N. Baliko, and S. Bowers, The relative playfulness of juvenile Lewis and Fischer-344 rats, Physiology & Behavior, vol.80, issue.2-3, pp.385-394, 1997.
DOI : 10.1016/j.physbeh.2003.09.002

E. F. Smith, Does play matter? Functional and evolutionary aspects of animal and human play, Behavioral and Brain Sciences, vol.88, issue.01, pp.139-155, 1991.
DOI : 10.1016/S0163-1047(80)90482-3

V. D. Son, Time budgets of Macaca fascicularis in a mangrove forest, pp.1-4, 2004.

M. Spinka, R. C. Newberry, and M. Bekoff, Mammalian Play: Training for the Unexpected, The Quarterly Review of Biology, vol.76, issue.2, pp.141-168, 2001.
DOI : 10.1086/393866

B. Thierry, Patterns of agonistic interactions in three species of macaque (Macaca mulatta, M. fascicularis, M. tonkeana) Aggressive Behavior, pp.223-233, 1985.

G. Thinès, Le jeu ? Le jeu chez l'enfant. Encyclopedia Universalis, pp.33-37, 1996.

N. Tinbergen, "Derived" Activities; Their Causation, Biological Significance, Origin, and Emancipation During Evolution, The Quarterly Review of Biology, vol.27, issue.1, pp.1-32, 1952.
DOI : 10.1086/398642

V. Trezza, P. J. Baarendse, and L. J. Vanderschuren, The pleasures of play: pharmacological insights into social reward mechanisms, Trends in Pharmacological Sciences, vol.31, issue.10, pp.463-469, 2010.
DOI : 10.1016/j.tips.2010.06.008

V. Trezza, R. Damsteegt, E. J. Achterberg, and L. J. Vanderschuren, Nucleus Accumbens ??-Opioid Receptors Mediate Social Reward, Journal of Neuroscience, vol.31, issue.17, pp.6362-6370, 2011.
DOI : 10.1523/JNEUROSCI.5492-10.2011

S. P. Turner, M. Ewen, J. A. Rooke, and S. A. Edwards, The effect of space allowance on performance, aggression and immune competence of growing pigs housed on straw deep-litter at different group sizes, Livestock Production Science, vol.66, issue.1, pp.47-55, 2000.
DOI : 10.1016/S0301-6226(00)00159-7

F. N. Van-hasselt, J. M. Tieskens, V. Trezza, H. J. Krugers, L. J. Vanderschuren et al., Within-litter variation in maternal care received by individual pups correlates with adolescent social play behavior in male rats, Physiology & Behavior, vol.106, issue.5, pp.701-706, 2012.
DOI : 10.1016/j.physbeh.2011.12.007

J. A. Van-hoof, The facial displays of the Catarrhine monkeys and apes, Primate ethology, pp.7-68, 1967.

I. Veissier, D. Lamy, and P. Le-neindre, Social behaviour in domestic beef cattle when yearling calves are left with the cows for the next calving, Applied Animal Behaviour Science, vol.27, issue.3, pp.193-200, 1990.
DOI : 10.1016/0168-1591(90)90056-J

G. Waring, Horse behavior, 2003.

S. M. Wells and B. Goldschmidt-rothschild, Social Behaviour and Relationships in a Herd of Camargue Horses, Zeitschrift f??r Tierpsychologie, vol.16, issue.6, pp.363-380, 1979.
DOI : 10.1111/j.1439-0310.1979.tb00299.x

M. West, Social Play in the Domestic Cat, American Zoologist, vol.14, issue.1, pp.427-436, 1974.
DOI : 10.1093/icb/14.1.427

A. Wolff and M. Hausberger, Behaviour of Foals before Weaning May Have Some Genetic Basis, Ethology, vol.49, issue.1, pp.1-10, 1994.
DOI : 10.1111/j.1439-0310.1994.tb00876.x

A. Wolff, M. Hausberger, and N. Le-scolan, Experimental tests to assess emotionality in horses, Behavioural Processes, vol.40, issue.3, pp.209-221, 1997.
DOI : 10.1016/S0376-6357(97)00784-5
URL : https://hal.archives-ouvertes.fr/hal-01319160

D. G. Wood-gush, K. Vestergaard, and H. V. Petersen, The significance of motivation and environment in the development of exploration in pigs, Biology Behaviour, vol.15, pp.39-45, 1990.

F. Xu, L. Xie, X. Li, Q. Li, T. Wang et al., Construction and Validation of a Systematic Ethogram of Macaca fascicularis in a Free Enclosure, PLoS ONE, vol.32, issue.Suppl 2, p.37486, 2012.
DOI : 10.1371/journal.pone.0037486.s003