G. Burke, O. Fiehn, and N. Moran, Effects of facultative symbionts and heat stress on the metabolome of pea aphids, The ISME Journal, vol.12, issue.2, pp.242-252, 2009.
DOI : 10.1016/0020-1790(80)90043-8

P. Cassey, T. Blackburn, R. Duncan, and S. Chown, Concerning invasive species: Reply to Brown and Sax, Austral Ecology, vol.21, issue.4, pp.475-480, 2005.
DOI : 10.1139/cjz-77-9-1337

M. Chevrier, Introduction de deux espèces d'insectes aux Iles Kerguelen: processus de colonisation et exemples d'interactions, Thèse de, 1996.

S. Chown, N. Gremmen, and K. Gaston, Ecological biogeography of Southern Ocean Islands: species?area relationships, human impacts, and conservation, Am Nat, vol.152, pp.562-575, 1998.

S. Chown, A. Hoffmann, T. Kristensen, M. Angiletta, N. Stenseth et al., Adapting to climate change: a perspective from evolutionary physiology, Climate Research, vol.43, issue.1, pp.3-15, 2010.
DOI : 10.3354/cr00879

S. Chown, A. Huiskes, N. Gremmen, J. Lee, A. Terauds et al., Continent-wide risk assessment for the establishment of nonindigenous species in Antarctica, Proceedings of the National Academy of Sciences, vol.109, issue.13, pp.4938-4943, 2012.
DOI : 10.1073/pnas.1119787109

URL : https://hal.archives-ouvertes.fr/hal-00696311

S. Chown and V. Smith, Climate change and the short-term impact of feral house mice at the sub-Antarctic Prince Edward Islands, Oecologia, vol.10, issue.4, pp.508-516, 1993.
DOI : 10.1007/BF00320508

H. Colinet, T. Hance, P. Vernon, A. Bouchereau, and D. Renault, Does fluctuating thermal regime trigger free amino acid production in the parasitic wasp Aphidius colemani (Hymenoptera: Aphidiinae)?, Comparative Biochemistry and Physiology Part A: Molecular & Integrative Physiology, vol.147, issue.2, pp.484-492, 2007.
DOI : 10.1016/j.cbpa.2007.01.030

URL : https://hal.archives-ouvertes.fr/hal-00145790

H. Colinet, V. Larvor, M. Laparie, and R. D. , Exploring the plastic response to cold acclimation through metabolomics, Functional Ecology, vol.105, issue.3, pp.711-722, 2012.
DOI : 10.1111/j.1365-2435.2012.01985.x

URL : https://hal.archives-ouvertes.fr/hal-00717742

P. Convey, R. Key, R. Key, M. Belchier, and C. Waller, Recent range expansions in non-native predatory beetles on sub-Antarctic South Georgia, Polar Biology, vol.8, issue.4, pp.597-602, 2011.
DOI : 10.1007/s00300-010-0909-6

C. Deutsch, J. Tewksbury, R. Huey, K. Sheldon, C. Ghalambor et al., Impacts of climate warming on terrestrial ectotherms across latitude, Proceedings of the National Academy of Sciences, vol.105, issue.18, pp.6668-6672, 2008.
DOI : 10.1073/pnas.0709472105

A. Elbein, Y. Pan, I. Pastuszak, and D. Carroll, New insights on trehalose: a multifunctional molecule, Glycobiology, vol.13, issue.4, pp.17-27, 2013.
DOI : 10.1093/glycob/cwg047

O. Fiehn, G. Wohlgemuth, M. Scholz, T. Kind, D. Lee et al., Quality control for plant metabolomics: reporting MSI-compliant studies, The Plant Journal, vol.138, issue.4, pp.691-704, 2008.
DOI : 10.1111/j.1365-313X.2007.03387.x

Y. Frenot, S. Chown, J. Whinam, P. Selkirk, P. Convey et al., Biological invasions in the Antarctic: extent, impacts and implications, Biological Reviews, vol.62, issue.207, pp.45-72, 2005.
DOI : 10.1016/S0169-5347(01)02194-2

Y. Frenot, J. Gloaguen, L. Massé, and M. Lebouvier, Human activities, ecosystem disturbance and plant invasions in subantarctic Crozet, Kerguelen and Amsterdam Islands, Biological Conservation, vol.101, issue.1, pp.33-50, 2001.
DOI : 10.1016/S0006-3207(01)00052-0

S. Gill and N. Tuteja, Polyamines and abiotic stress tolerance in plants, Plant Signaling & Behavior, vol.45, issue.1, pp.26-33, 2010.
DOI : 10.4161/psb.5.1.10291

S. Hazell and J. Bale, Low temperature thresholds: Are chill coma and CTmin synonymous?, Journal of Insect Physiology, vol.57, issue.8, pp.1085-1089, 2011.
DOI : 10.1016/j.jinsphys.2011.04.004

K. Hidalgo, M. Laparie, R. Bical, V. Larvor, A. Bouchereau et al., Metabolic fingerprinting of the responses to salinity in the invasive ground beetle Merizodus soledadinus at the Kerguelen Islands, Journal of Insect Physiology, vol.59, issue.1, pp.91-100, 2013.
DOI : 10.1016/j.jinsphys.2012.10.017

URL : https://hal.archives-ouvertes.fr/hal-00801308

R. Huey and J. Kingsolver, Evolution of thermal sensitivity of ectotherm performance, Trends in Ecology & Evolution, vol.4, issue.5, pp.131-135, 1989.
DOI : 10.1016/0169-5347(89)90211-5

P. Hulme, S. Bacher, M. Kenis, S. Klotz, I. Kühn et al., Grasping at the routes of biological invasions: a framework for integrating pathways into policy, Journal of Applied Ecology, vol.42, issue.2, pp.403-414, 2008.
DOI : 10.1111/j.1365-2664.2007.01442.x

D. Joanisse and K. Storey, Temperature acclimation and seasonal responses by enzymes in cold-hardy gall insects, Archives of Insect Biochemistry and Physiology, vol.131, issue.4, pp.339-349, 1995.
DOI : 10.1002/arch.940280404

P. Johns, Arthorpoda of the subantarctic islands of New Zealand (1) Coleoptera : Carabidae Southern New Zealand, Patagonian, and Falkland Islands Insular carabidae, Journal of the Royal Society of New Zealand, vol.27, issue.3, pp.283-302, 1974.
DOI : 10.1080/03036758.1974.10419396

S. Khodayari, S. Moharramipour, V. Larvor, K. Hidalgo, and R. D. , Deciphering the Metabolic Changes Associated with Diapause Syndrome and Cold Acclimation in the Two-Spotted Spider Mite Tetranychus urticae, PLoS ONE, vol.94, issue.1, p.54025, 2013.
DOI : 10.1371/journal.pone.0054025.s002

URL : https://hal.archives-ouvertes.fr/hal-00812070

V. Ko?tál, J. Vambera, and J. Bastl, On the nature of pre-freeze mortality in insects: water balance, ion homeostasis and energy charge in the adults of Pyrrhocoris apterus, Journal of Experimental Biology, vol.207, issue.9, pp.1509-1521, 2004.
DOI : 10.1242/jeb.00923

V. Ko?tál, R. D. Rozsypal, and J. , Seasonal changes of free amino acids and thermal hysteresis in overwintering heteropteran insect, Pyrrhocoris apterus, Comparative Biochemistry and Physiology Part A: Molecular & Integrative Physiology, vol.160, issue.2, pp.245-251, 2011.
DOI : 10.1016/j.cbpa.2011.06.017

N. Kruger and A. Von-schaewen, The oxidative pentose phosphate pathway: structure and organisation, Current Opinion in Plant Biology, vol.6, issue.3, pp.236-246, 2003.
DOI : 10.1016/S1369-5266(03)00039-6

L. Lalouette, V. Kostal, H. Colinet, D. Gagneul, and R. D. , Cold exposure and associated metabolic changes in adult tropical beetles exposed to fluctuating thermal regimes, FEBS Journal, vol.44, issue.7, pp.1759-1767, 2007.
DOI : 10.1111/j.1742-4658.2007.05723.x

URL : https://hal.archives-ouvertes.fr/hal-00154786

L. Lalouette, C. Williams, M. Cottin, B. Sinclair, and R. D. , Thermal biology of the alien ground beetle Merizodus soledadinus introduced to the Kerguelen Islands, Polar Biology, vol.6, issue.4, pp.509-517, 2012.
DOI : 10.1007/s00300-011-1096-9

URL : https://hal.archives-ouvertes.fr/hal-00688852

M. Laparie, V. Larvor, Y. Frenot, and R. D. , Starvation resistance and effects of diet on energy reserves in a predatory ground beetle (Merizodus soledadinus; Carabidae) invading the Kerguelen Islands, Comparative Biochemistry and Physiology Part A: Molecular & Integrative Physiology, vol.161, issue.2, pp.122-129, 2012.
DOI : 10.1016/j.cbpa.2011.09.011

URL : https://hal.archives-ouvertes.fr/hal-00677499

M. Laparie, R. Bical, V. Larvor, P. Vernon, Y. Frenot et al., Habitat phenotyping of two sub-Antarctic flies by metabolic fingerprinting: Evidence for a species outside its home?, Comparative Biochemistry and Physiology Part A: Molecular & Integrative Physiology, vol.162, issue.4, pp.406-412, 2012.
DOI : 10.1016/j.cbpa.2012.04.022

URL : https://hal.archives-ouvertes.fr/hal-00717402

M. Lebouvier, M. Laparie, M. Hullé, A. Marais, Y. Cozic et al., The significance of the sub-Antarctic Kerguelen Islands for the assessment of the vulnerability of native communities to climate change, alien insect invasions and plant viruses, Biological Invasions, vol.22, issue.5, pp.1195-1208, 2011.
DOI : 10.1007/s10530-011-9946-5

URL : https://hal.archives-ouvertes.fr/hal-01111159

F. Lefebvre and J. Fourche, Mitochondrial aspects of cold acclimation in Pieris brassicae (Lepidoptera) during diapause and post-diapause imaginal development, Journal of Thermal Biology, vol.10, issue.1, pp.29-33, 1985.
DOI : 10.1016/0306-4565(85)90007-5

E. Lombaert, T. Guillemaud, J. Cornuet, T. Malausa, B. Facon et al., Bridgehead Effect in the Worldwide Invasion of the Biocontrol Harlequin Ladybird, PLoS ONE, vol.5, issue.3, p.9743, 2010.
DOI : 10.1371/journal.pone.0009743.s007

A. Malmendal, J. Overgaard, J. Bundy, J. Sørensen, N. Nielsen et al., Metabolomic profiling of heat stress: hardening and recovery of homeostasis in Drosophila, AJP: Regulatory, Integrative and Comparative Physiology, vol.291, issue.1, pp.205-212, 2006.
DOI : 10.1152/ajpregu.00867.2005

W. Mamai, K. Mouline, C. Blais, V. Larvor, K. Dabiré et al., s.l. Mosquitoes Exposed to the Stressful Conditions of the Dry Season in Burkina Faso, West Africa, Physiological and Biochemical Zoology, vol.87, issue.3, pp.486-497, 2014.
DOI : 10.1086/675697

URL : https://hal.archives-ouvertes.fr/hal-01299974

D. Mcmullen and K. Storey, Mitochondria of cold hardy insects: Responses to cold and hypoxia assessed at enzymatic, mRNA and DNA levels, Insect Biochemistry and Molecular Biology, vol.38, issue.3, pp.367-373, 2008.
DOI : 10.1016/j.ibmb.2007.12.003

M. Michaud, J. Benoit, G. Lopez-martinez, M. Elnitsky, R. Lee et al., Metabolomics reveals unique and shared metabolic changes in response to heat shock, freezing and desiccation in the Antarctic midge, Belgica antarctica, Journal of Insect Physiology, vol.54, issue.4, pp.645-655, 2008.
DOI : 10.1016/j.jinsphys.2008.01.003

P. Ottesen, Diel activity patterns of Carabidae, Staphylinidae and Perimylopidae (Coleoptera) at South Georgia, Sub-Antarctic, Polar Biology, vol.10, issue.7, pp.515-519, 1990.
DOI : 10.1007/BF00233700

J. Overgaard, A. Malmendal, J. Sørensen, J. Bundy, V. Loeschcke et al., Metabolomic profiling of rapid cold hardening and cold shock in Drosophila melanogaster, Journal of Insect Physiology, vol.53, issue.12, pp.1218-1232, 2007.
DOI : 10.1016/j.jinsphys.2007.06.012

L. Petraccone, S. Baiano, G. Fiorentino, G. Barone, and C. Giancola, Simultaneous effect of cadaverine and osmolytes on ct-DNA thermal stability, Thermochimica Acta, vol.418, issue.1-2, pp.47-52, 2004.
DOI : 10.1016/j.tca.2003.11.052

R. Development and C. Team, R: A Language and Environment for Statistical Computing, pp.3-900051, 2008.

D. Rangel, A. Anderson, and D. Roberts, Evaluating physical and nutritional stress during mycelial growth as inducers of tolerance to heat and UV-B radiation in Metarhizium anisopliae conidia, Mycological Research, vol.112, issue.11, pp.1362-1372, 2008.
DOI : 10.1016/j.mycres.2008.04.013

G. Ruijter, M. Bax, H. Patel, S. Flitter, P. Van-de-vondervoort et al., Mannitol Is Required for Stress Tolerance in Aspergillus niger Conidiospores, Eukaryotic Cell, vol.2, issue.4, pp.690-698, 2003.
DOI : 10.1128/EC.2.4.690-698.2003

H. Rhee, E. Kim, and J. Lee, Physiological polyamines: simple primordial stress molecules, Journal of Cellular and Molecular Medicine, vol.168, issue.4, pp.685-703, 2007.
DOI : 10.1104/pp.009951

S. Roig-junent and M. Dominguez, Diversidad de la familia Carabidae (Coleoptera) en Chile, Revista chilena de historia natural, vol.74, issue.3, pp.549-571, 2001.
DOI : 10.4067/S0716-078X2001000300006

M. Salvucci, Sorbitol accumulation in whiteflies: evidence for a role in protecting proteins during heat stress, Journal of Thermal Biology, vol.25, issue.5, pp.353-361, 2000.
DOI : 10.1016/S0306-4565(99)00107-2

D. Siaussat, M. Laparie, A. Maria, and R. D. , Heat shock protein responses to salinity, food deprivation, and temperature in the invasive ground beetle Merizodus soledadinus at the Kerguelen Islands, Polar Biology, vol.150, issue.2, pp.201-209, 2013.
DOI : 10.1007/s00300-012-1252-x

URL : https://hal.archives-ouvertes.fr/hal-00797891

D. Simberloff, How common are invasion-induced ecosystem impacts?, Biological Invasions, vol.9, issue.Supplement 1, pp.1255-1268, 2011.
DOI : 10.1007/s10530-011-9956-3

J. Storey and K. Storey, Cold hardiness and freeze tolerance Functional metabolism: regulation and adaption, pp.473-503, 2005.
DOI : 10.1002/047167558x.ch17

C. Todd, Respiratory metabolism in two species of carabid beetle from the sub-Antarctic island of South Georgia, Polar Biology, vol.18, issue.3, pp.66-171, 1997.
DOI : 10.1007/s003000050173

C. Todd and W. Block, Responses to desiccation in four Coleopterans from sub-Antarctic South Georgia, Journal of Insect Physiology, vol.43, issue.10, pp.905-913, 1997.
DOI : 10.1016/S0022-1910(97)00055-3

P. Vitousek, H. Mooney, J. Lubchenco, and J. Melillo, Human Domination of Earth's Ecosystems, Science, vol.277, issue.5325, pp.494-499, 1997.
DOI : 10.1126/science.277.5325.494

J. Yoder, J. Benoit, D. Denlinger, and D. Rivers, Stress-induced accumulation of glycerol in the flesh fly, Sarcophaga bullata: Evidence indicating anti-desiccant and cryoprotectant functions of this polyol and a role for the brain in coordinating the response, Journal of Insect Physiology, vol.52, issue.2, pp.202-214, 2006.
DOI : 10.1016/j.jinsphys.2005.10.005

M. Waller, R. Whitney, M. Taylor, H. Miyagawa, K. Matsuda et al., Multiresidue pesticides analysis using synchronous scan/SIM mode GC?MS, LC GC N Am, vol.25, p.39, 2007.