T. Aavik, R. Holderegger, and J. Bolliger, The structural and functional connectivity of the 684 grassland plant Lychnis flos-cuculi, Heredity, vol.112, pp.471-478, 2014.

D. Adriaens, O. Honnay, and M. Hermy, No evidence of a plant extinction debt in highly 686 fragmented calcareous grasslands in Belgium, Biol Conserv, vol.133, pp.212-224, 2006.

F. Adriaensen, J. P. Chardon, D. Blust, G. Swinnen, E. Villalba et al., , p.689, 2003.

, The application of 'leastcost' modelling as a functional landscape model, Landsc Urban Plan, vol.690, pp.242-248

T. Allen and T. B. Starr, Hierarchy: Perspectives for Ecological Complexity, p.692, 1988.

E. Andersson and C. Nilsson, Temporal variation in the drift of plant litter and propagules in 694 a small boreal river, Freshw Biol, vol.47, pp.1674-1684, 2002.

C. Andreasen, H. Stryhn, and J. C. Streibig, Decline of the Flora in Danish Arable Fields, J Appl 696 Ecol, vol.33, pp.619-626, 1996.

S. Archer and D. A. Pyke, Plant-animal interactions affecting plant establishment and 698 persistence on revegetated rangeland, J Range Manag, vol.44, pp.558-565, 1991.

A. Arellano-rivas, J. A. De-nova, and M. Ma, Patch isolation and shape predict 700 plant functional diversity in a naturally fragmented forest, J Plant Ecol, pp.1-11, 2016.

A. G. Auffret, Y. Rico, J. M. Bullock, D. Hooftman, R. J. Pakeman et al.,

A. Traveset, H. Wagner, and S. Cousins, Plant functional connectivity -integrating 704 landscape structure and effective dispersal, J Ecol, 2017.

C. Ayram, M. E. Mendoza, A. Etter, and D. Salicrup, Habitat connectivity in biodiversity 706 conservation: A review of recent studies and applications, Prog Phys Geogr Earth Environ, vol.707, pp.7-37, 2016.

C. Baessler and S. Klotz, Effects of changes in agricultural land-use on landscape structure 709 and arable weed vegetation over the last 50 years, Agric Ecosyst Environ, vol.115, pp.43-50, 2006.

P. Beier and R. F. Noss, Do Habitat Corridors Provide Connectivity, Conserv Biol, vol.12, pp.1241-712, 1998.

A. F. Bennett, Linkages in the Landscape: The Role of Corridors and Connectivity in 714, 2003.

, Wildlife Conservation. IUCN-The World Conservation Union

A. J. Bennett, J. Q. Radford, and A. Haslem, Properties of Land Mosaics: Implications for Nature 716 Conservation in Agricultural Environments, Biol Conserv, vol.133, pp.250-264, 2006.

J. Betbeder, L. Hubert-moy, F. Burel, S. Corgne, and J. Baudry, Assessing ecological habitat 719 structure from local to landscape scales using synthetic aperture radar, Ecol Indic, vol.52, pp.545-557, 2015.

J. Betbeder, J. Nabucet, E. Pottier, J. Baudry, S. Corgne et al., Detection and 722 Characterization of Hedgerows Using TerraSAR-X Imagery, Remote Sens, vol.6, pp.3752-3769, 2014.

G. Boedeltje, J. P. Bakker, R. M. Bekker, J. M. Van-groenendael, and M. Soesbergen, Plant 725 dispersal in a lowland stream in relation to occurrence and three specific life-history traits of 726 the species in the species pool, J Ecol, vol.91, pp.855-866, 2003.

G. Boedeltje, J. P. Bakker, T. Brinke, A. Van-groenendael, J. M. Soesbergen et al., Dispersal 728 phenology of hydrochorous plants in relation to discharge, seed release time and buoyancy of 729 seeds: the flood pulse concept supported, J Ecol, vol.92, pp.786-796, 2004.

G. Bornette, C. Amoros, and N. Lamouroux, Aquatic plant diversity in riverine wetlands: the 732 role of connectivity, Freshw Biol, vol.39, pp.267-283, 1998.

D. R. Bowne and M. A. Bowers, Interpatch movements in spatially structured populations: a 734 literature review, Landsc Ecol, vol.19, 2004.

S. V. Brückmann, J. Krauss, and I. Steffan-dewenter, Butterfly and plant specialists suffer from 736 reduced connectivity in fragmented landscapes, J Appl Ecol, vol.47, pp.799-809, 2010.

L. A. Brudvig, Interpreting the effects of landscape connectivity on community diversity, 2016.

, J Veg Sci, vol.27, pp.4-5

L. A. Brudvig, E. I. Damschen, J. J. Tewksbury, N. M. Haddad, and D. J. Levey, , p.Landscape, 2009.

, connectivity promotes plant biodiversity spillover into non-target habitats, Proc Natl Acad Sci, vol.742, pp.9328-9332

F. Burel and J. Baudry, Ecologie du paysage. Concepts, méthodes et applications. Editions 744 TEC et DOC, 1999.
URL : https://hal.archives-ouvertes.fr/hal-01210029

J. M. Calabrese and W. F. Fagan, A comparison-shopper's guide to connectivity metrics, 2004.
DOI : 10.2307/3868383

, Ecol Environ, vol.2, pp.529-536

C. Chetkiewicz, C. Clair, and M. S. Boyce, Corridors for Conservation: Integrating 748 Pattern and Process, Annu Rev Ecol Evol Syst, vol.37, pp.317-342, 2006.
DOI : 10.1146/annurev.ecolsys.37.091305.110050

J. Cornelissen, S. Lavorel, E. Garnier, S. Díaz, N. Buchmann et al., , p.751

H. Steege, H. D. Morgan, M. Van-der-heijden, J. G. Pausas, and H. Poorter, A handbook of 752 protocols for standardised and easy measurement of plant functional traits worldwide, Aust J 753 Bot, vol.51, pp.335-380, 2003.

S. Cousins, H. Ohlson, and O. Eriksson, Effects of historical and present fragmentation on 755 plant species diversity in semi-natural grasslands in Swedish rural landscapes, Landsc Ecol, vol.756, 2007.

S. Cousins and D. Vanhoenacker, Detection of extinction debt depends on scale and 758 specialisation, Biol Conserv, vol.144, pp.782-787, 2011.

E. I. Damschen, D. V. Baker, G. Bohrer, R. Nathan, J. L. Orrock et al., How fragmentation and corridors affect wind dynamics 761 and seed dispersal in open habitats, Proc Natl Acad Sci, vol.111, pp.3484-3489, 2014.

E. I. Damschen, L. A. Brudvig, N. M. Haddad, D. J. Levey, J. L. Orrock et al., The 764 movement ecology and dynamics of plant communities in fragmented landscapes, Proc Natl, vol.765, 2008.

, Acad Sci, vol.105, pp.19078-19083

E. I. Damschen, N. M. Haddad, J. L. Orrock, J. J. Tewksbury, and D. J. Levey, Corridors increase 767 plant species richness at large scales, Science, vol.313, pp.1284-1286, 2006.
DOI : 10.1126/science.1130098

D. De-ryck, E. Robert, N. Schmitz, T. Van-der-stocken, D. Nitto et al., , p.769

N. Koedam, Size does matter, but not only size: Two alternative dispersal strategies for 770 viviparous mangrove propagules, Aquat Bot, vol.103, pp.66-73, 2012.

J. M. Diamond, Biogeographic kinetics: estimation of relaxation times for avifaunas of 772 southwest pacific islands, Proc Natl Acad Sci, vol.69, pp.3199-3203, 1972.

Z. Dzwonko and S. Loster, Species richness of small woodlands on the western Carpathian 774 foothills, vol.76, pp.15-27, 1988.

O. Eriksson, Regional Dynamics of Plants: A Review of Evidence for Remnant, p.776, 1996.

, Sink and Metapopulations. Oikos, vol.77, pp.248-258

M. Evju, S. Blumentrath, O. Skarpaas, and O. E. Stabbetorp, Plant species occurrence in a 778 fragmented grassland landscape: the importance of species traits, Biodivers Conserv, vol.24, pp.547-779, 2015.

M. A. Leibold, M. Holyoak, N. Mouquet, P. Amarasekare, J. M. Chase et al., The metacommunity concept: a 881 framework for multi-scale community ecology, Ecol Lett, vol.7, pp.601-613, 2004.

D. J. Levey, B. M. Bolker, J. J. Tewksbury, S. Sargent, and N. M. Haddad, Effects of landscape 884 corridors on seed dispersal by birds, Science, vol.309, pp.146-148, 2005.

R. Lindborg, Evaluating the distribution of plant life-history traits in relation to current 886 and historical landscape configurations, J Ecol, vol.95, pp.555-564, 2007.

R. Lindborg and O. Eriksson, Historical Landscape Connectivity Affects Present Plant 889, 2004.
DOI : 10.1890/04-0367

, Species Diversity. Ecology, vol.85, pp.1840-1845

J. P. Lindgren and S. Cousins, Island biogeography theory outweighs habitat amount 891 hypothesis in predicting plant species richness in small grassland remnants, Landsc Ecol, vol.892, pp.1895-1906, 2017.

J. B. Logue, N. Mouquet, H. Peter, and H. Hillebrand, Empirical approaches to 894 metacommunities: a review and comparison with theory, Trends Ecol Evol, vol.26, pp.482-491, 2011.
DOI : 10.1016/j.tree.2011.04.009

M. Luoto, S. Rekolainen, J. Aakkula, and J. Pykälä, Loss of Plant Species Richness and Habitat 897, 2003.

, Connectivity in Grasslands Associated with Agricultural Change in Finland, AMBIO J Hum, vol.898, pp.447-452

R. Macarthur and R. Levins, The Limiting Similarity, Convergence, and Divergence of 900, 1967.

, Coexisting Species, Am Nat, vol.101, pp.377-385

R. H. Macarthur and E. Wilson, The Theory of Island Biogeography, p.902, 1967.

. Press,

B. H. Mcrae, B. G. Dickson, T. H. Keitt, and V. B. Shah, Using Circuit Theory to Model 904, 2008.

, Connectivity in Ecology, Evolution, and Conservation. Ecology, vol.89, pp.2712-2724

E. S. Minor and R. H. Gardner, Landscape connectivity and seed dispersal characteristics inform 907 the best management strategy for exotic plants, Ecol Appl Publ Ecol Soc Am, vol.21, pp.739-749, 2011.

E. S. Minor, S. M. Tessel, K. Engelhardt, and T. R. Lookingbill, The role of landscape 909 connectivity in assembling exotic plant communities: a network analysis, Ecology, vol.90, pp.1802-910, 2009.

H. L. Moggridge, A. M. Gurnell, and J. O. Mountford, Propagule input, transport and deposition 912 in riparian environments: the importance of connectivity for diversity, J Veg Sci, vol.20, 2009.

A. Moilanen and M. Nieminen, Simple Connectivity Measures in Spatial Ecology, Ecology, vol.915, pp.1131-1145, 2002.

R. G. Morato, B. Ferraz-kmpm-de, R. C. De-paula, and C. B. De-campos, , 2014.

, Conservation Areas and Potential Corridors for Jaguars in the Caatinga Biome, Brazil. PLoS, vol.918

M. A. Mouchet, S. Villéger, N. Mason, and D. Mouillot, Functional diversity measures: an 920 overview of their redundancy and their ability to discriminate community assembly rules, Funct 921 Ecol, vol.24, pp.867-876, 2010.

N. Mouquet, E. Miller, T. Daufresne, T. , M. Kneitel et al., Consequences of varying regional 923 heterogeneity in source-sink metacommunities, Oikos, vol.113, pp.481-488, 2006.

N. Mouquet and M. Loreau, Community patterns in source-sink metacommunities, Am Nat, vol.926, pp.544-557, 2003.

H. T. Murphy and J. Lovett-doust, Context and connectivity in plant metapopulations and 928 landscape mosaics: does the matrix matter?, Oikos, vol.105, pp.3-14, 2004.

T. Naaf and J. Kolk, Colonization credit of post-agricultural forest patches in NE Germany 931 remains 130-230years after reforestation, Biol Conserv, vol.182, pp.155-163, 2015.

R. Nathan, Long-distance dispersal of plants, Science, vol.313, pp.786-788, 2006.

R. Nathan, G. G. Katul, H. S. Horn, S. M. Thomas, R. Oren et al., , p.936, 2002.

, Mechanisms of long-distance dispersal of seeds by wind, Nature, vol.418, pp.409-413

C. Nilsson, R. L. Brown, R. Jansson, and D. M. Merritt, The role of hydrochory in structuring 939 riparian and wetland vegetation, Biol Rev Camb Philos Soc, vol.85, pp.837-858, 2010.

R. V. O'neill, D. L. Deangelis, J. B. Waide, and T. Allen, , p.942, 1986.

, Ecosystems

J. L. Orrock and E. I. Damschen, Corridors Cause Differential Seed Predation, Ecol Appl, vol.944, pp.793-798, 2005.

J. L. Orrock, B. J. Danielson, M. J. Burns, and D. J. Levey, Spatial Ecology of Predator-Prey, vol.946, 2003.

, Interactions: Corridors and Patch Shape Influence Seed Predation, Ecology, vol.84, pp.2589-2599

O. Ovaskainen and I. Hanski, Transient Dynamics in Metapopulation Response to, 2002.

, Perturbation. Theor Popul Biol, vol.61, pp.285-295

S. W. Pacala and D. Tilman, Limiting similarity in mechanistic and spatial models of plant 951 competition in heterogeneous environments, Am Nat, vol.143, pp.222-257, 1994.

M. Pärtel, R. Szava-kovats, and M. Zobel, Dark diversity: shedding light on absent species, 2011.

, Trends Ecol Evol, vol.26, pp.124-128

L. Pascual-hortal and S. Saura, Comparison and development of new graph-based landscape 955 connectivity indices: towards the priorization of habitat patches and corridors for conservation, 2006.

, Landsc Ecol, vol.21, pp.959-967

B. D. Patterson and W. Atmar, Nested subsets and the structure of insular mammalian faunas 958 and archipelagos, Biol J Linn Soc, vol.96, pp.65-82, 1986.

K. Piessens, O. Honnay, and M. Hermy, The role of fragment area and isolation in the 960 conservation of heathland species, Biol Conserv, vol.122, pp.61-69, 2005.

K. Piessens, O. Honnay, K. Nackaerts, and M. Hermy, Plant Species Richness and Composition 962 of Heathland Relics in North-Western Belgium: Evidence for a Rescue-Effect?, J Biogeogr, vol.963, pp.1683-1692, 2004.

B. Pollux, A. Luteijn, J. M. Van-groenendael, and N. J. Ouborg, Gene flow and genetic 965 structure of the aquatic macrophyte Sparganium emersum in a linear unidirectional river, 2009.

, Freshw Biol, vol.54, pp.64-76

D. A. Pyke and S. Archer, Plant-plant interactions affecting plant establishment and persistence 968 on revegetated rangeland, J Range Manag, vol.44, pp.550-557, 1991.

B. Rayfield, M. Fortin, and A. Fall, The sensitivity of least-cost habitat graphs to relative cost 970 surface values, Landsc Ecol, vol.25, pp.519-532, 2010.

K. Riibak, T. Reitalu, R. Tamme, A. Helm, P. Gerhold et al., , p.972

H. C. Prentice and M. Pärtel, Dark diversity in dry calcareous grasslands is determined by 973 dispersal ability and stress-tolerance, Ecography, vol.38, pp.713-721, 2015.

M. R. Rossetti, T. Tscharntke, R. Aguilar, and P. Batáry, Responses of insect herbivores and 975 herbivory to habitat fragmentation: a hierarchical meta-analysis, Ecol Lett, vol.20, pp.264-272, 2017.

U. G. Sandström, Green Infrastructure Planning in Urban Sweden, Plan Pract Res, vol.17, pp.373-978, 2002.

S. Saura and L. Pascual-hortal, A new habitat availability index to integrate connectivity in 980 landscape conservation planning: Comparison with existing indices and application to a case 981 study, Landsc Urban Plan, vol.83, pp.91-103, 2007.

M. Scanlan, Biogeography of forest plants in the Prairie-Forest Ecotone in Western 983, 1981.

. Minnesota, Forest Island Dynamics in Man-dominated 984

. Landscapes and . Springer, , pp.97-124

A. Schleicher, R. Biedermann, and M. Kleyer, Dispersal traits determine plant response to 986 habitat connectivity in an urban landscape, Landsc Ecol, vol.26, pp.529-540, 2011.
DOI : 10.1007/s10980-011-9579-1

D. Simberloff and J. Cox, Consequences and Costs of Conservation Corridors, Conserv Biol, vol.989, issue.1, pp.63-71, 1987.

H. Soomers, D. N. Winkel, Y. Du, and M. J. Wassen, The dispersal and deposition of 991 hydrochorous plant seeds in drainage ditches, Freshw Biol, vol.55, pp.2032-2046, 2010.

A. E. Sorensen, Seed Dispersal by Adhesion, Annu Rev Ecol Syst, vol.17, pp.443-463, 1986.

S. F. Spear, N. Balkenhol, M. Fortin, B. H. Mcrae, and K. Scribner, Use of resistance surfaces 996 for landscape genetic studies: considerations for parameterization and analysis, Mol Ecol, vol.997, pp.3576-3591, 2010.

C. Stoate, N. Boatman, R. Borralho, C. R. Carvalho, G. R. De-snoo et al., Ecological 999 impacts of arable intensification in Europe, J Environ Manage, vol.63, pp.337-365, 2001.

J. Stöcklin and E. Winkler, Optimum reproduction and dispersal strategies of a clonal plant 1002 in a metapopulation: a simulation study with Hieracium pilosella, Evol Ecol, vol.18, pp.563-584, 2004.

A. Suárez-esteban, M. Delibes, and J. M. Fedriani, Barriers or corridors? The overlooked role 1005 of unpaved roads in endozoochorous seed dispersal, J Appl Ecol, vol.50, pp.767-774, 2013.

L. L. Sullivan, B. L. Johnson, L. A. Brudvig, and N. M. Haddad, Can dispersal mode predict corridor 1008 effects on plant parasites?, Ecology, vol.92, pp.1559-1564, 2011.
DOI : 10.1890/10-1116.1

URL : https://esajournals.onlinelibrary.wiley.com/doi/pdf/10.1890/10-1116.1

O. Tackenberg, Modeling Long-Distance Dispersal of Plant Diaspores by Wind, Ecol 1010 Monogr, vol.73, pp.173-189, 2003.
DOI : 10.1890/0012-9615(2003)073[0173:mldopd]2.0.co;2

P. D. Taylor, L. Fahrig, and K. A. With, Landscape connectivity: a return to the basics, p.1012, 2006.
DOI : 10.1017/cbo9780511754821.003

, Connectivity Conservation: maintaining connections for nature, p.1013

. Cambridge, , pp.29-43

P. D. Taylor, L. Fahrig, K. Henein, and G. Merriam, Connectivity Is a Vital Element of 1015, 1993.

, Landscape Structure. Oikos, vol.68, pp.571-573

J. J. Tewksbury, D. J. Levey, N. M. Haddad, S. Sargent, J. L. Orrock et al., , p.1017

J. Brinkerhoff, E. I. Damschen, and P. Townsend, Corridors affect plants, animals, and their 1018 interactions in fragmented landscapes, Proc Natl Acad Sci, vol.99, pp.12923-12926, 2002.

J. Thiele, S. Buchholz, and J. Schirmel, Using resistance distance from circuit theory to model 1021 dispersal through habitat corridors, J Plant Ecol rtx004, 2017.

D. Tilman, R. M. May, C. L. Lehman, and M. A. Nowak, Habitat destruction and the extinction 1023 debt, Nature, vol.371, pp.65-66, 1994.

L. Tischendorf and L. Fahrig, On the usage and measurement of landscape connectivity, 2000.

, Oikos, vol.90, pp.7-19

T. Tscharntke, J. M. Tylianakis, T. A. Rand, R. K. Didham, L. Fahrig et al., , p.1027

Y. , C. To, C. F. Dormann, R. M. Ewers, J. Fründ et al., , p.1028

C. Kremen, D. A. Landis, L. W. Lindenmayer, D. Scherber, C. Sodhi et al.,

I. , T. C. Van-der-putten, W. H. Westphal, and C. , Landscape moderation of biodiversity 1030 patterns and processes -eight hypotheses, Biol Rev, vol.87, pp.661-685, 2012.

K. Tzoulas, K. Korpela, S. Venn, V. Yli-pelkonen, A. Ka?mierczak et al., , p.1033, 2007.

, Promoting ecosystem and human health in urban areas using Green Infrastructure: A literature 1034 review, Landsc Urban Plan, vol.81, pp.167-178

A. G. Van-der-valk, Succession in Wetlands: A Gleasonian Approach, Ecology, vol.62, pp.688-1036, 1981.

M. Vellend, J. A. Myers, S. Gardescu, and P. L. Marks, Dispersal of Trillium seeds by deer: 1038 implications for long-distance migration of forest herbs, Ecology, vol.84, pp.1067-1072, 2003.

K. Verheyen, M. Vellend, H. V. Calster, G. Peterken, and M. Hermy, Metapopulation dynamics 1041 in changing landscapes: a new spatially realistic model for forest plants, Ecology, vol.85, pp.3302-1042, 2004.

C. Violle, M. Navas, D. Vile, E. Kazakou, C. Fortunel et al., Let the 1044 concept of trait be functional! Oikos, vol.116, pp.882-892, 2007.

P. Vittoz and R. Engler, Seed dispersal distances: a typology based on dispersal modes and 1046 plant traits, Bot Helvetica, vol.117, pp.109-124, 2007.

E. Weiher and P. A. Keddy, The Assembly of Experimental Wetland Plant Communities, 1995.

, Oikos, vol.73, pp.323-335

D. S. Wilson, Complex Interactions in Metacommunities, with Implications for 1050, 1992.

, Biodiversity and Higher Levels of Selection. Ecology, vol.73, pp.1984-2000

E. O. Wilson and E. O. Willis, Applied Biogeography, p.1052, 1975.

, Ecology and Evolution of Communities, pp.522-534

D. Zacharias and D. Brandes, Species Area-Relationships and Frequency: Floristical Data 1054, 1990.
DOI : 10.1007/bf00032600

URL : https://publikationsserver.tu-braunschweig.de/servlets/MCRFileNodeServlet/dbbs_derivate_00003113/fulltext.pdf

, Analysis of 44 Isolated Woods in Northwestern Germany, Vegetatio, vol.88, pp.21-29

K. A. Zeller, K. Mcgarigal, and A. R. Whiteley, Estimating landscape resistance to movement: 1057 a review, Landsc Ecol, vol.27, pp.777-797, 2012.

, we only refer to connected patches versus unconnected patches. resulting from the logistic regression between patch occupancy for each species 1132 as explained by connectivity intensity), species richness in the different 1133 categories of the seed longevity index, proportion of species characterised by a 1134 trait, trait value of response groups to connectivity and species trait values 1135 d. Community weighted mean traits and functional diversity approach: functional 1136 gamma diversity (standard effect size values calculated for Rao's quadratic 1137 entropy traducing functional divergence or convergence) and gamma aggregated 1138 mean trait values (gamma scale weighed mean trait values, p.1140

, Concerning actual dispersal approaches, we included studies at the species level because very 1142 few studies have assessed the actual dispersal of communities in relation to connectivity, vol.1143, p.1144