G. Walther, Ecological responses to recent climate change, Nature, vol.416, pp.389-395, 2002.

S. L. Chown and C. J. Klok, Altitudinal body size clines: latitudinal effects associated with changing seasonality, Ecography, vol.26, pp.445-455, 2003.

R. Hickling, The distributions of a wide range of taxonomic groups are expanding polewards, Glob Change Biol, vol.12, pp.450-455, 2006.

D. Renault, M. Laparie, S. J. Mccauley, and D. Bonte, Environmental adaptations, ecological filtering and dispersal, central to insect invasions, Annu Rev Entomol, vol.63, pp.345-368, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01696977

C. Robinet and A. Roques, Direct impacts of recent climate warming on insect populations, Integr Zool, vol.5, pp.132-142, 2010.

J. J. Hellmann, J. E. Byers, B. G. Bierwagen, and J. S. Dukes, Five potential consequences of climate change for invasive species, Conserv Biol, vol.22, pp.534-543, 2008.

E. L. Charnov and J. F. Gillooly, Thermal time: body size, food quality and the 10 degrees C rule, Evol Ecol Res, vol.5, pp.43-51, 2003.

K. A. Hughes, S. Ott, M. Bölter, and P. Convey, (eds) Trends in Antarctic terrestrial and limnetic ecosystems: Antarctica as a global indicator, pp.35-54, 2006.

M. Williamson and A. Fitter, The varying success of invaders, Ecology, vol.77, pp.1661-1666, 1996.

Y. Frenot, Biological invasions in the Antarctic: extent, impacts and implications, Biol Rev, vol.80, pp.45-72, 2005.

J. J. Stachowicz, J. R. Terwin, R. B. Whitlatch, and R. W. Osman, Linking climate change and biological invasions: Ocean warming facilitates nonindigenous species invasions, Proc Natl Acad Sci, vol.99, pp.15497-15500, 2002.

K. B. Suttle, M. A. Thomsen, and M. E. Power, Species interactions reverse grassland responses to changing climate, Science, vol.315, pp.640-642, 2007.

M. Vilà, How well do we understand the impacts of alien species on ecosystem services? A pan-European, cross-taxa assessment, Front Ecol Environ, vol.8, pp.135-144, 2010.

N. L. Ward and G. J. Masters, Linking climate change and species invasion: an illustration using insect herbivores, Glob Change Biol, vol.13, pp.1605-1615, 2007.

Y. Frenot, J. Gloaguen, and P. Trehen, Climate change in Kerguelen Islands and colonization of recently deglaciated areas by Poa kerguelensis and P. annua, Antarctic communities: species, structure and survival, pp.358-366

M. Ingimarsdóttir, J. Ripa, Ó. B. Magnúsdóttir, and K. Hedlund, Food web assembly in isolated habitats: A study from recently emerged nunataks, Basic Appl Ecol, vol.14, pp.174-183, 2013.

V. R. Smith and M. Steenkamp, Climatic change and its ecological implications at a subantarctic island, Oecologia, vol.85, pp.14-24, 1990.

D. M. Bergstrom and S. L. Chown, Life at the front: history, ecology and change on Southern Ocean Islands, Trends Ecol Evol, vol.14, pp.472-477, 1999.

M. Greve, R. Mathakutha, C. Steyn, and S. L. Chown, Terrestrial invasions on sub-Antarctic Marion and Prince Edward Islands, Bothalia, vol.47, 2017.

M. Lebouvier, The significance of the sub-Antarctic Kerguelen Islands for the assessment of the vulnerability of native communities to climate change, alien insect invasions and plant viruses, Biol Invasions, vol.13, pp.1195-1208, 2011.
URL : https://hal.archives-ouvertes.fr/hal-01111159

M. Laparie and D. Renault, Physiological responses to temperature in Merizodus soledadinus (Col., Carabidae), a subpolar carabid beetle invading sub-Antarctic islands, Polar Biol, vol.39, pp.35-45, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01255879

R. Jeannel, Les Trechides de la Paléantarctide occidentale, pp.527-655, 1962.

P. Roux and J. F. Voisin, Notes sur les Carabiques des iles Falkland (Col. Caraboidea), Bull -Soc Entomol Fr, pp.200-204, 1982.

M. Chevrier, P. Vernon, and Y. Frenot, Potential effects of two alien insects on a subantarctic wingless fly in the Kerguelen Islands, Antarctic communities: species, structure and survival, pp.424-431, 1997.

T. Ouisse, M. Laparie, M. Lebouvier, and D. Renault, New insights into the ecology of Merizodus soledadinus, a predatory carabid beetle invading the sub-Antarctic Kerguelen Islands, Polar Biol, vol.40, pp.2201-2209, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01654276

S. L. Chown, N. J. Gremmen, and K. J. Gaston, Ecological biogeography of Southern Ocean Islands: species-area relationships, human impacts, and conservation, Am Nat, vol.152, pp.562-575, 1998.

L. Lalouette, Thermal biology of the alien ground beetle Merizodus soledadinus introduced to the Kerguelen Islands, Polar Biol, vol.35, pp.509-517, 2012.
URL : https://hal.archives-ouvertes.fr/hal-00688852

T. Ouisse, The importance of relative humidity and trophic resources in governing ecological niche of the invasive carabid beetle Merizodus soledadinus in the Kerguelen archipelago, J Insect Physiol, pp.42-49, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01371825

D. Renault, Characterization of the habitats colonized by the alien ground beetle Merizodus soledadinus at the Kerguelen islands, Rev Ecol, vol.70, pp.28-32, 2015.
URL : https://hal.archives-ouvertes.fr/hal-02391384

R. I. Leihy, G. A. Duffy, E. Nortje, and S. L. Chown, Data Descriptor: High resolution temperature data for ecological research and management on the Southern Ocean Islands, Scientific Data, vol.5, 2018.

M. E. Dillon, M. R. Frazier, and R. Dudley, Into thin air: physiology and evolution of alpine insects, Integr Comp Biol, vol.46, pp.49-61, 2006.

J. F. Jenkin, . Macquarie-island, and . Subantarctic, (eds) Structure and function of tundra ecosystems, Ecological Bulletins, pp.375-397, 1975.

E. A. Hugo-coetzee and P. C. Le-roux, Distribution of microarthropods across altitude and aspect in the sub-Antarctic: climate change implications for an isolated oceanic island, Acarologia, vol.53, pp.43-60, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01883375

M. Hullé, D. Pannetier, J. C. Simon, P. Vernon, and Y. Frenot, Aphids of sub-Antarctic Iles Crozet and Kerguelen: species diversity, host range and spatial distribution, Antarctic Science, vol.15, pp.203-209, 2003.

K. F. Davies, B. A. Melbourne, J. L. Mcclenahan, and T. Tuff, Statistical models for monitoring and predicting effects of climate change and invasion on the free-living insects and a spider from sub-Antarctic Heard Island, Polar Biol, vol.34, pp.119-125, 2011.

K. F. Davies and B. A. Melbourne, Statistical models of invertebrate distribution on Macquarie Island: a tool to assess climate change and local human impacts, Polar Biol, vol.21, pp.240-250, 1999.

N. J. Gremmen, S. L. Chown, and D. J. Marshall, Impact of the introduced grass Agrostis stolonifera on vegetation and soil fauna communities at Marion Island, sub-Antarctic, Biol Conserv, vol.85, pp.223-231, 1998.

C. K. Mirth and L. M. Riddiford, Size assessment and growth control: how adult size is determined in insects, BioEssays, vol.29, pp.344-355, 2007.

A. P. Moczek, Horn polyphenism in the beetle Onthophagus taurus: larval diet quality and plasticity in parental investment determine adult body size and male horn morphology, Behav Ecol, vol.9, pp.636-641, 1998.

L. Partridge and V. French, Thermal evolution of ectotherm body size: why get big in the cold, Animals and Temperature: Phenotypic and Evolutionary Adaptation, pp.265-292

, , 1996.

B. Ciplak, D. Sirin, M. S. Taylan, and S. Kaya, Altitudinal size clines, species richness and population density: case studies in Orthoptera, J Orthoptera Res, vol.17, pp.157-163, 2008.

B. Krasnov, D. Ward, and G. Shenbrot, Body size and leg length variation in several species of darkling beetles (Coleoptera: Tenebrionidae) along a rainfall and altitudinal gradient in the Negev Desert (Israel), J Arid Environ, vol.34, pp.477-489, 1996.

T. Sota, Altitudinal variation in life cycles of carabid beetles: life-cycle strategy and colonization in alpine zones, Arctic and Alpine Research, vol.28, pp.441-447, 1996.

T. M. Van-der-have and G. De-jong, Adult size in ectotherms: temperature effects on growth and differentiation, J Theor Biol, vol.183, pp.329-340, 1996.

D. M. O'brien, M. L. Fogel, and C. L. Boggs, Renewable and nonrenewable resources: Amino acid turnover and allocation to reproduction in Lepidoptera, Proc Natl Acad Sci, vol.99, pp.4413-4418, 2002.

N. Tigreros, Linking nutrition and sexual selection across life stages in a model butterfly system, Funct Ecol, vol.27, pp.145-154, 2013.

D. E. Wheeler, I. Tuchinskaya, N. A. Buck, and B. E. Tabashnik, Hexameric storage proteins during metamorphosis and egg production in the diamondback moth, Plutella xylostella (Lepidoptera), J Insect Physiol, vol.46, issue.99, pp.202-206, 2000.

N. M. Teets and D. L. Denlinger, Physiological mechanisms of seasonal and rapid cold-hardening in insects, Physiol Entomol, vol.38, pp.105-116, 2013.

P. H. Yancey, Organic osmolytes as compatible, metabolic and counteracting cytoprotectants in high osmolarity and other stresses, J Exp Biol, vol.208, pp.2819-2830, 2005.

K. Hidalgo, Metabolic fingerprinting of the responses to salinity in the invasive ground beetle Merizodus soledadinus at the Kerguelen Islands, J Insect Physiol, vol.59, pp.91-100, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00801308

H. Colinet, T. Hance, and P. Vernon, Water relations, fat reserves, survival, and longevity of a cold-exposed parasitic wasp Aphidius colemani (Hymenoptera: Aphidiinae), Environ Entomol, vol.35, pp.228-236, 2006.
URL : https://hal.archives-ouvertes.fr/hal-00076619

L. Lalouette, Cold exposure and associated metabolic changes in adult tropical beetles exposed to fluctuating thermal regimes, FEBS J, vol.274, pp.1759-1767, 2007.
URL : https://hal.archives-ouvertes.fr/hal-00141865

V. Foray, A handbook for uncovering the complete energetic budget in insects: the van Handel's method (1985) revisited, Physiol Entomol, vol.37, pp.295-302, 2012.

M. Laparie, V. Larvor, Y. Frenot, and D. Renault, Starvation resistance and effects of diet on energy reserves in a predatory ground beetle
URL : https://hal.archives-ouvertes.fr/hal-00677499

, Carabidae) invading the Kerguelen Islands, Comp Biochem Physiol A Mol Integr Physiol, vol.161, pp.122-129, 2012.

J. G. Bundy, M. P. Davey, and M. R. Viant, Environmental metabolomics: a critical review and future perspectives, Metabolomics, vol.5, pp.3-21, 2009.

M. R. Viant, Recent developments in environmental metabolomics, Mol Biosyst, vol.4, 2008.

S. Khodayari, Deciphering the metabolic changes associated with diapause syndrome and cold acclimation in the two-spotted spider mite Tetranychus urticae, PloS ONE, vol.8, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00812070

K. Mangor, N. K. Drønen, K. H. Kaergaard, and S. E. Kristensen, Shoreline Management Guidelines. DHI, p.450, 2017.

M. Robin, J. L. Chapuis, and M. Lebouvier, Remote sensing of vegetation cover change in islands of the Kerguelen archipelago, Polar Biol, vol.34, pp.1689-1700, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00666745

P. Breheny and W. Burchett, visreg: Visualization ofregression models

S. Lê, J. Josse, and F. Husson, FactoMineR: an R package for multivariate analysis, J Stat Softw, vol.25, pp.1-18, 2008.

W. N. Venables and B. D. Ripley, Random and mixed effects, Modern Applied Statistics with S, pp.271-300, 2002.

, Core Team R: A language and environment for statistical computing. R Foundation for Statistical Computing, 2016.