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M. R. , M. D. , W. H. Conceived-the-study, D. B. , S. F. et al., wrote the manuscript with input from all coauthors. Competing interests: The authors declare that they have no competing interests. Data and materials availability: All data needed to evaluate the conclusions in the paper are present in the paper and/or the Supplementary Materials. Individual bam files (aligned human reads) are available through the European Nucleotide Archive under accession number PRJEB36208, performed the laboratory work. M.R. performed the data analyses, 2019.

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