C. B. Kasper, V. Bastazini, J. Salvi, and C. Hamilton, Trophic ecology and the use of 1031 shelters and latrines by the Neotropical otter Lontra longicaudis in the Taquari Valley, p.1032, 2008.

. Southern-brazil-iheringia, Zoologia, vol.98, pp.469-474

R. W. Kays and J. L. Gittleman, The social organization of the kinkajou Potos flavus 1034 (Procyonidae), Journal of Zoology, vol.253, pp.491-504, 2001.

R. W. Kays, J. L. Gittleman, and R. K. Wayne, Microsatellite analysis of kinkajou social 1036 organization, Mol Ecol, vol.9, pp.743-751, 2000.

B. Keane, P. M. Waser, S. R. Creel, N. M. Creel, L. F. Elliott et al., Subordinate 1038 reproduction in dwarf mongooses, Anim Behav, vol.47, pp.65-75, 1994.

L. Keller, Indiscriminate altruism, unduly nice parents and siblings, TREE, vol.12, pp.99-103, 1997.

W. A. Khan, M. Qasim, A. E. Chaudhry, A. A. Bhaagat, H. B. Akhtar et al., , 2010.

C. Otter, Lutrogale perspicillata sindica. in Pakistan, Pak J Zool, vol.42, pp.817-824

W. D. Koenig, F. A. Pitelka, W. J. Carmen, R. L. Mumme, and M. T. Stanback, The evolution of delayed 1043 dispersal in cooperative breeders, Q Rev Biol, vol.67, pp.111-150, 1992.

K. P. Koepfli, K. A. Deere, G. J. Slater, C. Begg, K. Begg et al., , 1045.

. Rk, Multigene phylogeny of the Mustelidae, Resolving relationships, tempo and 1046 biogeographic history of a mammalian adaptive radiation, B M C Biol, vol.6, p.10, 2008.

K. P. Koepfli and R. K. Wayne, Phylogenetic relationships of otters. Carnivora, Mustelidae. 1048 based on mitochondrial cytochrome B sequences, J Zool Lond, vol.246, pp.401-416, 1998.

H. Kokko and J. Ekman, Delayed dispersal as a route to breeding; territorial inheritance, 'safe 1050 havens' and ecological constraints, Am Nat, vol.160, pp.468-484, 2002.

H. Kokko and R. A. Johnstone, Social queuing in animal societies. a dynamic model of 1052 reproductive skew, Proc R Soc Lond B Biol Sc, vol.266, pp.571-578, 1999.

J. Komdeur, Importance of habitat saturation and territory quality for evolution of 1054 cooperative breeding in the seychelles warbler, Nature, vol.358, pp.493-495, 1992.

J. Komdeur, Influence of helping and breeding experience on reproductive performance in 1056 the Seychelles warbler. A translocation experiment, Behav Ecol, vol.7, pp.326-333, 1996.

J. Komdeur, Variation in individual investment strategies among social animals, Ethology, vol.1058, pp.729-747, 2006.

J. Krause and G. D. Ruxton, Living in groups, 2002.

M. Krkosek, J. S. Lauzon-guay, and M. A. Lewis, Relating dispersal and range expansion of 1061, 2007.

, California sea otters, Theor Pop Biol, vol.71, pp.401-407

H. Kruuk, Wild Otters, predation and populations, 1995.

, Kruuk H (2006) Otters. ecology, behavior and conservation

H. Kruuk and A. Moorehouse, The spatial organization of otters Lutra lutra in Shetland, J Zool, vol.224, pp.41-57, 1065.

H. Kruuk, B. Kanchanasaka, S. O'sullivan, and S. Wanghongsa, Niche separation in 3 sympatric 1067 otters lutra-perspicillata, l-lutra Aonyx cinerea in Huai-Kha-Khaeng, Biol Conserv, vol.69, pp.115-120, 1068.

H. Kruuk and P. C. Goudswaard, Effects of changes in fish populations in Lake Victoria on the 1070 food of otters Lutra maculicollis Schinz and Aonyx capensis Lichtenstein, Afr J Zool, vol.28, pp.322-1071, 1990.

Z. Ladds, W. Hoppitt, and N. J. Boogert, Social learning in otters, R Soc open sci, vol.30, pp.4-8, 2017.

K. L. Laidre, R. J. Jameson, E. Gurarie, S. J. Jeffries, and H. Alllen, Spatial habitat use patterns of 1075 sea otters in coastal Washington, J Mamm, vol.90, pp.906-917, 2009.

S. Larivière, Lontra provocax, Mammal Sp, vol.610, pp.1-4, 1999.

S. Larivière, Aonyx congicus. Mammal Sp, vol.650, pp.1-3, 2001.

S. Larivière, Amblonyx cinereus. Mammal Sp, vol.720, pp.1-5, 2003.

S. Larivière and L. R. Walton, Lontra canadensis, Mammal Sp, vol.587, pp.1-8, 1998.

L. Galliard, J. F. Rémy, A. Ims, R. A. Lambin, and X. , Patterns and processes of dispersal 1081 behaviour in arvicoline rodents, Mol Ecol, vol.21, pp.505-523, 2012.

A. Lejeune, Ethologie des loutres Hydrictis maculicollis au lac Muhazi, 1989.

, Mammalia, vol.53, pp.191-202

A. Lemasson, R. A. Palombit, and R. Jubin, Friendships between males and lactating females in 1085 a free-ranging group of olive baboons (Papio hamadryas anubis). Evidence for playback 1086 experiments, Behav Ecol Sociobiol, vol.62, pp.1027-1035, 2008.

A. Lemasson, M. A. Mikus, C. Blois-heulin, and T. Lodé, Social partner discrimination based on 1088 sounds and scents in Asian small-clawed otters (Aonyx cinereus), Naturwissenschaft, vol.100, pp.1089-275, 2013.

C. Leuchtenberger and G. Mourão, Social organization and territoriality of giant otters, 2008.

. Carnivora, Mustelidae in a seasonally flooded savanna in Brazil, Sociobiology, vol.52, pp.257-270

B. M. Leuthold, Social organization and behaviour of giraffe in Tsavo East National Park, 1979.

, Afr J Ecol, vol.17, pp.19-34

J. D. Ligon and S. H. Ligon, Reciprocity in the green woodhoopoe Phoeniculus purpureu, Anim 1095 Behav, vol.31, pp.480-489, 1983.

T. Lodé, Conspecific tolerance and sexual segregation in the use of space and habitats 1097 in the European polecat, Acta Ther, vol.41, pp.171-176, 1996.

T. Lodé, Polecat predation on frogs and toads at breeding sites in western France, Ethol, vol.1099, 1996.

, Ecol Evol, vol.8, pp.115-124

T. Lodé, Trophic status and feeding habits of the European Polecat Mustela putorius L., 1101 1758, Mammal Review, vol.27, pp.177-184, 1997.

T. Lodé, Functional response and area-restricted search of a predator, seasonal 1103 exploitation of anurans by European polecat Mustela putorius, Austral Ecol, vol.25, pp.223-231, 2000.

T. Lodé, V. Pereboom, and R. Berzins, Implications of an individualistic lifestyle for species 1111 conservation, lessons from jealous beasts, C R Biol, vol.326, pp.30-36, 2003.

D. W. Luca and J. R. Ginsberg, Dominance, reproduction and survival in banded mongooses, 1113 towards an egalitarian social system?, Anim Behav, vol.61, pp.17-30, 2001.

G. X. Ludwig, V. Hokka, R. Sulkava, and H. Ylönen, Otter Lutra lutra. predation on farmed and 1115 free-living salmonids in boreal freshwater habitats, Wildl Biol, vol.8, pp.203-208, 2002.

D. W. Macdonald, The ecology of carnivore social behavior, Nature, vol.301, pp.379-384, 1983.

D. W. Macdonald and C. Sillero-zubiri, The biology and conservation of wild canids, 1118.

D. Macdonald and C. Newman, Musteloid sociality: the grass-roots of society, 2018.

D. Newman, C. Lauren, and A. H. , Biology and conservation of musteloid, pp.167-188

L. J. Macshane, J. A. Estes, M. L. Riedman, and M. M. Staedler, Repertoire, structure and individual 1122 variation of vocalizations in the sea otter, J Mam, vol.76, pp.414-427, 1995.

J. C. Mangel, T. Whitty, M. Vogel, G. , A. Shigueto et al., , p.1124, 2010.

, Latitudinal variation in diet and patterns of human interaction in the marine otter, 1125.

, Sc, vol.27, pp.14-25

M. B. Manser, Response of foraging group members to sentinel calls in suricates, Suricata 1127 suricatta, Proc R Soc. B, vol.266, pp.1013-1019, 1999.

M. B. Manser, The acoustic structure of suricates' alarm calls varies with predator type and 1129 the level of response urgency, Proc R Soc B, vol.268, pp.2315-2324, 2001.

C. F. Mason and S. M. Macdonald, Otters -ecology and conservation, p.1131, 1986.

. Cambridge,

M. M. Mbizah, M. Valeix, D. W. Macdonald, and A. J. Loveridge, Applying the resource dispersion 1133 hypothesis to a fission-fusion society: A case study of the African lion, p.1134, 2019.

. Evol, , vol.9, pp.9111-9119

D. L. Mech and L. Boitani, Wolves, behavior, ecology and conservation, p.1136, 2003.

. Chicago,

G. Medina-vogel and C. Gonzalez-lagos, Habitat use and diet of endangered southern river 1138, 2008.

, otter Lontra provocax in a predominantly palustrine wetland in Chile, Wildl Biol, vol.14, pp.211-220

G. Medina-vogel, J. L. Bartheld, R. A. Pacheco, and R. Delgado, Population assessment and 1140 habitat use of marine otter Lontra felina in southern Chile, J 1143 Mammal, vol.12, pp.487-494, 2006.

G. Medina-vogel, C. Delgado, and R. E. Alvarez, Feeding ecology of the marine otter Lutra felina 1145 in a rocky seashore of the south of Chile, Mar Mamm Sci, vol.20, pp.134-144, 2004.

W. E. Melquist and M. G. Hornocker, Ecology of river otters in west central Idaho, Wildl Monogr, vol.83, pp.1-60, 1147.

J. C. Mitani, D. A. Merriwether, and C. Zhang, Male affiliation, cooperation and kinship in wild 1149 chimpanzees, Anim Behav, vol.59, pp.885-893, 2000.

L. M. Möller, L. B. Beheregaray, R. G. Harcourt, and M. Krutzen, Alliance membership and kinship 1151 in wild male bottlenose dolphins (Tursiops aduncus) of southeastern Australia, Proc R Soc B 1152, vol.268, pp.1941-1947, 2001.

D. H. Monson, J. A. Estes, J. L. Bodkin, and D. B. Siniff, Life history plasticity and population 1154 regulation in sea otters, Oikos, vol.90, pp.457-468, 2000.

A. Mosser and C. Packer, Group territoriality and the benefits of sociality in the African lion, p.1156, 2009.

P. Leo, Anim Behav, vol.78, pp.359-370

D. C. Newman and C. R. Griffin, Wetland use by river otters in Massachusetts, J Wildl Manag, vol.58, pp.1158-1176, 1994.

X. D. Nguyen, Current status of otters Mammalia Lutrinae in Viet Nam with conservation 1160 implications, Tiger Paper, vol.33, pp.8-14, 2005.

X. D. Nguyen, T. A. Pham, and H. T. Le, New Information about the Hairy-Nosed Otter Lutra 1162 sumatrana in Vietnam, IUCN Otter Sp Group Bull, vol.18, pp.64-75, 2001.

P. Nonacs, Measuring and using skew in the study of social behavior and evolution, Am, vol.1164, pp.577-589, 2000.

S. Okasha, Genetic relatedness and the evolution of altruism, Phil Science, vol.69, pp.138-149, 2002.

R. Ostfeld, L. Ebensperger, L. Klosterman, and J. Castilla, Foraging, activity budget, and social 1167 behavior of the South American marine otter Lutra feline, Nat Geog Res, vol.5, pp.422-438, 1989.

C. Packer, Reciprocal altruism in Papio Anubis, Nature, vol.265, pp.441-443, 1977.

C. Packer and L. Ruttan, The evolution of cooperative hunting, Am Nat, vol.132, pp.159-198, 1988.

C. Packer, D. A. Gilbert, A. E. Pusey, O. 'brien, and S. J. , A molecular genetic analysis of kinship and 1171 cooperation in African lions, Nature, vol.351, pp.562-565, 1991.

S. Pagacz and J. Witczuk, Intensive exploitation of amphibians by Eurasian otter Lutra lutra in 1173 the Wolosaty stream, southern Poland, Ann Zool Fenn, vol.47, pp.403-410, 2010.

D. M. Parker, R. K. Burchell, and R. Bernard, The diet of Cape clawless otters at two sites 1175 along the Bloukrans River, Af Zool, vol.40, pp.330-334, 2005.

R. Pardini, Feeding Ecology of the Neotropical River Otter Lontra longicaudis in an 1177, 1998.

, Atlantic Forest Stream, south-eastern Brazil, J Zool, vol.245, pp.385-391

, Alaska. Aquatic Mammals, vol.31, pp.226-233

S. M. Pellis, Some Aspects of Play-Fighting in a Captive Group of Oriental Small-Clawed 1181, 1984.

A. Otters and . Cinerea, Z Tierpsychol (Ethology), vol.65, pp.77-83

E. Pennisi, How did cooperative behavior evolve, Science, vol.309, p.93558, 2005.

B. M. Perdue, R. J. Snyder, and T. L. Maple, Cognitive Research in Asian Small-Clawed Otters. Int 1184, J Comp Psychol, vol.26, pp.105-113, 2013.

A. Perinchery, Conservation genetics of Indian otters and their habitat use in Eravikulam 1186 National Park, 2008.

A. Perinchery, D. Jathanna, and A. Kumar, Factors determining occupancy and habitat use by 1188, 2011.

, Asian small-clawed otters in the Western Ghats, India. J Mammal, vol.92, pp.796-802

M. R. Perrin, D. Carranza, and I. , Habitat use by spotted-necked otters in the KwaZulu-1190, 2000.

, Natal Drakensberg. Afr J Wildl Res, vol.30, pp.8-14

N. Perrin and V. Mazalov, Local competition, inbreeding and the evolution of sex-biased 1192 dispersal, Am Nat, vol.155, pp.116-127, 2000.

T. R. Pope, The reproductive consequences of male cooperation in the red howler monkey, 1194 paternity exclusion in multi-male and single-male troops using genetic markers, Behav Ecol 1195 Sociobiol, vol.27, pp.439-446, 1990.

R. A. Powell, Mustelid spacing patterns, variations on a theme by Mustela, Z Tierpsychol, vol.50, pp.153-165, 1197.

N. Prakash, Factors influencing the occurrence and habitat use of small-clawed otters 1199 (Aonyx cinerea) in the human-modified landscape of Western Ghats, p.1200, 2010.

, Nat. Cent. Biol. Sciences

C. Prigioni, A. Balestrieri, L. Remonti, A. Gargaro, and G. Priore, Diet of the Eurasian otter Lutra 1202 lutra. in relation to freshwater habitats and alien fish species in southern Italy, Ethol Ecol Evol, vol.18, pp.307-320, 2006.

L. M. Prox and D. R. Farine, A framework for conceptualizing dimensions of social organization 1205 in mammals, Ecology & Evolution, vol.10, pp.791-807, 2020.

A. E. Pusey, Sex-biased dispersal and inbreeding avoidance in birds and mammals, TREE 1207, vol.2, pp.295-299, 1987.

L. Quaglietta, V. Fonseca-da-silva, M. A. Boitani, and L. , , p.1209, 2014.

, Eurasian otter ( Lutra lutra ), J Mamm, vol.95

A. Raha and S. A. Hussain, Factors affecting habitat selection by three sympatric otter species 1212 in the southern Western Ghats, India. Acta Ecol Sin, vol.36, pp.45-49, 2016.

,

S. Ratnayeke, G. A. Tuskan, and M. R. Pelton, Genetic relatedness and female spatial 1215 organization in a solitary carnivore, the raccoon, Procyon lotor, paludinosus. Af J Ecol, vol.11, pp.237-253, 1216.

H. K. Reeve and L. Keller, Partitioning of reproduction in mother-daughter versus sibling 1219 associations, a test of optimal skew theory, Am Nat, vol.145, pp.119-132, 1995.

J. Reiczigel, Z. Lang, L. Rózsa, and B. Tóthmérész, Measures of sociality, two different views of 1221 group size, Anim Behav, vol.75, pp.715-721, 2008.

D. G. Reid, T. E. Code, A. Reid, and S. M. Herrero, Food habits of the river otter in a boreal 1223 ecosystem, Can J Zool, vol.72, pp.1306-1313, 1994.

L. Remonti, C. Prigioni, A. Balestrieri, S. Sgrosso, and G. Priore, Trophic flexibility of the otter 1225 (Lutra lutra) in southern Italy, Mamm Biol, vol.73, pp.293-302, 2008.

E. Revilla, Moving beyond the resource dispersion hypothesis, TREE, vol.18, p.380, 2003.

E. Revilla and F. Palomares, Spatial organization, group living and ecological correlates in 1228 low-density populations of Eurasian badgers, Meles meles, J Anim Ecol, vol.71, pp.497-512, 2002.

M. L. Rheingantz, H. F. Waldemarin, L. Rodrigues, and T. P. Multon, Seasonal and spatial 1230 differences in feeding habits of the Neotropical otter Lontra longicaudis, Carnivora, p.1231, 2011.

, Mustelidae. in a coastal catchment of southeastern, Brazil Zool, vol.28, pp.37-44

C. Ribas, H. Cunha, G. Damasceno, W. Magnusson, A. Solé-cava et al., More than 1233 meets the eye: kinship and social organization in giant otters, p.1234, 2015.

, Ecol Sociobiol, vol.70

M. L. Riedman and J. A. Estes, The sea otter (Enhydra lutris): behavior, ecology, and natural 1236 history, U.S. Fish and Wildl Serv Biol Report, vol.90, issue.14, 1990.

K. R. Rock, E. S. Rock, R. T. Bowyer, and J. B. Faro, Degree of association and use of a helper by 1238 coastal river otters, Lutra canadensis, vol.108, pp.1239-367, 1994.

F. Rosas, J. Zuanon, and S. K. Carter, Feeding Ecology of the Giant Otter, Pteronura 1241 brasiliensis, vol.31, pp.502-506, 1999.

D. T. Rowe-rowe and M. J. Somers, Diet, foraging behaviour and coexistence of African otters 1243 and the water mongoose, Symp Zool Soc Lond, vol.71, pp.216-227, 1998.

D. R. Rubenstein and I. J. Lovette, Temporal environmental variability drives the evolution of 1245 cooperative breeding in birds, Curr Biol, vol.17, pp.1414-1419, 2007.

K. E. Ruckstuhl and P. Neuhaus, Sexual segregation in ungulates, vol.137, pp.361-377, 1247.

J. L. Sachs, The Evolution of Cooperation, Quart Rev Biol, vol.79, pp.135-160, 2004.

C. Schradin and N. Pillay, Intraspecific variation in the spatial and social organization of the 1250 African striped mouse, J Mammal, vol.86, pp.99-107, 2005.

R. Schröpfer, P. Wiegand, and H. H. Hogrefe, The implications of territoriality for the social 1252 system of the European pine marten Martes martes (L. 1758), Z Säugetierk, vol.62, pp.209-218, 1997.

M. A. Sepúlveda, J. L. Bartheld, R. Monsalve, V. Gómez, and G. Medina-vogel, Habitat use and 1254 spatial behaviour of the endangered Southern river otter Lontra provocax in riparian habitats 1255 of Chile, conservation implications, Biol Conserv, vol.140, pp.329-338, 2007.

M. A. Sepúlveda, J. L. Bartheld, C. Meynard, C. Astorga, M. Benavides et al., Landscape features and crustacean prey as predictors of the Southern river otter 1258 (Lontra provocax) distribution in Chile, Anim Conserv, vol.12, pp.522-530, 2009.

J. S. Shannon, Social organization and behavioral ontogeny of otters Lutra Canadensis in a 1260 coastal habitat in northern California, Bull Otter Specialist Group, vol.4, pp.8-13, 1991.

M. Schmelz, S. Duguid, M. Bohn, and C. Volter, Cooperative problem solving in giant otters 1262 (Pteronura brasiliensis) and Asian small-clawed otters (Aonyx cinerea), Anim Cogn, vol.20, p.1107, 2017.

,

P. W. Sherman, Nepotism and the evolution of alarm calls, Science, vol.197, pp.1246-1253, 1977.

L. Shorey, S. Piertney, J. Stone, and J. Hoglund, Fine scale genetic structuring on Manacus 1266 manacus leks, Nature, vol.408, pp.352-353, 2000.

J. B. Silk, The adaptive value of sociality in mammalian groups, Phil. Trans. R. Soc. B-Biol, 2007.

. Sci, , vol.362, pp.539-559

N. Sivasothi, The Small-clawed otter, Amblonyx cinereus (Illiger, 1815) (Carnivora, 1996.

. Mustelidae and . Lutrinae, Report of a family 1271 group on Pulau Tekong Besar and of the fate of wildlife there, The Pangolin, vol.7, pp.23-31

N. Sivasothi and B. Nor, A review of otters (Carnivora. Mustelidae. Lutrinae) in Malaysia 1273 and Singapore, Hydrobiologia, vol.285, pp.151-170, 1994.

T. Sjöåsen, Movements and establishment of reintroduced European otters Lutra lutra, J, vol.1275, 1997.

, App Ecol, vol.34, pp.1070-1080

J. E. Smith, J. M. Kolowski, K. E. Graham, S. E. Dawes, and K. E. Holekamp, Social and ecological 1277 determinants of fission-fusion dynamics in the spotted hyaena, Anim Behav, vol.76, pp.619-636, 2008.

J. E. Smith, E. M. Swanson, D. Reed, and K. E. Holekamp, , p.1279, 2012.

, Mammalian Carnivores and Its Relevance to Hominin Evolution, Curr Anthropol, vol.53, pp.436-452

C. T. Snowdon and K. A. Cronin, Cooperative breeders do cooperate, Behav Proc, vol.76, pp.138-141, 2007.

M. J. Somers, Foraging behaviour of Cape clawless otters (Aonyx capensis) in a marine 1282 habitat, J Zool Lond, vol.252, pp.473-480, 2000.

M. J. Somers and J. Nel, Diet in relation to prey of Cape clawless otters in two rivers in the 1284, 2003.

, Western Cape Province, South Africa, Afr J Ecol, vol.38, pp.317-326

G. Spong, J. Stone, S. Creel, and M. Bjorklund, Genetic structure of lions Panthera leo L, p.1286, 2002.

, Selous Game Reserve, implications for the evolution of sociality, J Evol Biol, vol.15, pp.945-953

K. A. Stiver, J. Fitzpatrick, J. K. Desjardins, and S. Balshine, Sex differences in rates of territory 1288 joining and inheritance in a cooperatively breeding cichlid fish, Anim Behav, vol.71, pp.449-456, 2006.

S. W. Townsend, M. Zöttl, and M. B. Manser, All clear? Meerkats attend to contextual information 1290 in close calls to coordinate vigilance, Behav Ecol Sociobiol, vol.65, pp.1927-1934, 2011.

P. Tyack, Population biology, social behavior and communication in whales and dolphins, vol.1293, pp.144-150, 1986.

J. Valqui, A review of its present status and 1295 implications for future conservation, Mammal Biol (Z Säugetierk), vol.77, pp.75-83, 2011.

S. L. Vehrencamp, Optimal degree of skew in cooperative societies, Am Zool, vol.23, pp.327-335, 1983.

J. L. Verdolin, Gunnison's prairie dog Cynomys gunnisoni testing the resource dispersion 1298 hypothesis, Behav Ecol Sociobiol, vol.63, pp.789-799, 2009.

G. Véron, M. Colyn, A. Dunham, P. Taylor, and P. Gaubert, Molecular systematics and origin of 1300 sociality in mongooses Herpestidae Carnivora, Mol Phyl Evol, vol.30, pp.582-598, 2004.

M. J. Villegas, A. A. Ebensperger, and L. A. , The influence of wave exposure on the foraging 1302 activity of marine otter, Lontra felina (Molina, 1782) (Carnivora Mustelidae) in northern Chile, 2007.

, J Ethol, vol.25, pp.281-286

A. P. Wagner, S. Creel, L. G. Frank, and S. T. Kalinowski, Patterns of relatedness and parentage in 1305 an asocial, polyandrous striped hyena population, Mol Ecol, vol.16, pp.4356-4369, 2007.

P. M. Waser, B. Keane, S. R. Creel, L. G. Elliot, and D. J. Minchella, Possible male coalitions in a 1307 solitary mongoose, Anim Behav, vol.47, pp.289-294, 1994.

S. K. Wasser and D. P. Barash, Reproductive suppression among female mammals, implications 1309 for biomedicine and sexual selection theory, Quart Rev Biol, vol.58, pp.513-538, 1983.

J. M. Weber, Seasonal exploitation of amphibians by otters Lutra lutra in north-east 1311, 1990.

. Scotland, J Zool, vol.220, pp.641-651

S. A. West, A. S. Griffin, and A. Gardner, Evolutionary explanations for cooperation, Curr Biol, vol.17, pp.1313-661, 2007.

S. A. West, I. Pen, and A. S. Griffin, Cooperation and competition between relatives, Science, vol.296, pp.1315-72, 2002.

W. Wickler and U. Seibt, Pedogenetic sociogenesis via the 'siblingroute' and consequences 1317 for Stegodyphus spiders, Ethology, vol.95, pp.1-18, 1993.

D. A. Williams and K. N. Rabenold, Male-biased dispersal, female philopatry and routes to 1319 fitness in a social corvid, J Anim Ecol, vol.74, pp.150-159, 2005.

D. S. Wilson, Weak Altruism, Strong Group Selection, Oikos, vol.59, pp.135-148, 1990.

R. W. Wrangham and D. I. Rubenstein, Social evolution in birds and mammals, Rubenstein 1322, 1986.

, Ecological aspects of social evolution in birds and mammals

J. Wright, H. Berg, D. Kort, S. R. Khazin, V. Maklakov et al., Cooperative sentinel behaviour in 1325 the Arabian babbler, Anim Behav, vol.62, pp.973-979, 2001.

J. Yamagiwa, J. Kahekwa, K. Basabose, and A. , Intra-specific variation in social 1327 organization of gorillas. implications for their social evolution, Primates, vol.44, pp.359-369, 2003.

, Reproductive Tactics of Free-Ranging American Minks, Mustela vison, J. Mammal, vol.85, pp.1330-432

H. Ylönen and J. S. Brown, Fear and the Foraging, Breeding and Sociality in Rodents, p.1332, 2007.

A. J. Young, A. A. Carlson, S. L. Monfort, A. F. Russell, and N. C. Bennett, Stress 1334 and the suppression of subordinate reproduction in cooperatively breeding meerkats, PNAS 1335, vol.103, pp.12005-12010, 2006.